/
primarieswhenpresentedwitharedteststimulus.Similarly, primarieswhenpresentedwitharedteststimulus.Similarly,

primarieswhenpresentedwitharedteststimulus.Similarly, - PDF document

briana-ranney
briana-ranney . @briana-ranney
Follow
421 views
Uploaded On 2016-06-12

primarieswhenpresentedwitharedteststimulus.Similarly, - PPT Presentation

intheuniqueconditionsubjectsassignedratingsofaboutfive toeachoftheredandyellowprimarieswhenpresentedwith anorangeteststimulusIfredgreenblueandyellowreally canbedistinguishedasuniqueweshouldexpec ID: 359773

intheuniquecondition subjectsassignedratingsofaboutfive toeachoftheredandyellowprimarieswhenpresentedwith anorangeteststimulus.Ifred green blue andyellowreally canbedistinguishedasunique weshouldexpec

Share:

Link:

Embed:

Download Presentation from below link

Download Pdf The PPT/PDF document "primarieswhenpresentedwitharedteststimul..." is the property of its rightful owner. Permission is granted to download and print the materials on this web site for personal, non-commercial use only, and to display it on your personal computer provided you do not modify the materials and that you retain all copyright notices contained in the materials. By downloading content from our website, you accept the terms of this agreement.


Presentation Transcript

primarieswhenpresentedwitharedteststimulus.Similarly, intheuniquecondition,subjectsassignedratingsofaboutfive toeachoftheredandyellowprimarieswhenpresentedwith anorangeteststimulus.Ifred,green,blue,andyellowreally canbedistinguishedasunique,weshouldexpectthatwhen theyarepresentedasteststimuliintheintermediatecondition theywouldbegivennear-zeroratings[Fig. 1(b) ].Thiswas clearlynotevidentintheresults. Isitpossiblethatindividualdifferencesinuniquehuesaf- fectourresults?Individualsvaryreliablyintheirsettingsof uniquehues[ 21 , 30 , 32 ],though within-subject variabilityin huesettingsaccountsforasubstantialportionofthetotalvari- abilityobserved[ 29 ].Weusedgroupmeanuniqueandinter- mediatehueselectionsasprimariesforourhuescalingtask. ThepredictionsshowninFig. 1(a) assumedthattheprimaries providedareunique,butthismightnotbetrueforparticular observerswhohaveuniquehuesettingsdifferentfromthe mean.If,foraparticularobserver,theprimariesprovided inthe “ unique ” conditionwerenotunique,wewouldexpect thepatternofresultstoresemblemorethepredictionforin- termediateprimariesshowninFig. 1(b) .Sinceresultsforthe majorityofobserversinbothconditionsmatchedthepredic- tionsforuniqueprimaries[Fig. 1(a) ],individualvariabilityin huesettingscannotundermineourconclusionthatthetypical patternofhuescalingresultsreportedintheliteraturecanbe achievedwithnon-uniqueprimaries. SternheimandBoynton[ 15 ]assessedfromtheirhuescaling resultswhetheragivenhuewasuniquebyapplyingthreecri- teria:(a)thatthecolorcategoryshouldbeusedreliably; (b)thattheratingfunctionforaprimaryshouldreachamaxi- muminaregionwherefunctionsforneighboringprimaries areataminimum;and(c)thataparticularuniquehueshould notberepresentedinratingsassignedtootherprimaries whentheprimaryassociatedwiththatuniquehueisnotper- mitted.Alltheprimariesweusedinbothuniqueandinter- mediateconditionssatisfiedSternheimandBoynton ’ s[ 15 ] firsttwocriteria(seeFig. 3 ).Takethefunctionfororange asanexample:thecategorywasusedreliablyasevidenced bysmallconfidenceintervalsandconsensusacrosssubjects, andthefunctionreachedamaximuminaregionwherefunc- tionsforneighboringhues(violetandlime)wereatamini- mum.Noprimariesfromeitherconditionfulfilled SternheimandBoynton ’ s[ 15 ]thirdcriterion.Theprimaries “ red, ”“ green, ”“ blue, ” and “ yellow ” werenotpermittedin theintermediatecondition.Yetwhenred,green,blue,oryel- lowteststimuliwerepresented,subjectswereabletomake responsesusingtheintermediateprimariestheywerepro- videdwith. “ Unique ” yellow,forexample,wasdescribedby assigningratingsofaboutfivetoeachofthepermittedprimar- iesoforangeandlime. Aminorityofcolorscientistshavequestionedtheassumed specialstatusofuniquehues.Jameson[ 32 ]hasdonesoby consideringthevariableresponsesofparticipantsinthe WorldColorSurvey[ 33 ].ApaperbySaundersandvanBrakel [ 34 ]questioningthelinguistic,neurophysiological,andpsy- chophysicalevidenceforuniquehuestriggeredmanyener- geticresponses.Theywereaccusedof “ throwingthebaby outwiththebathwater ” [ 35 ]and “ selectivevision ” [ 36 ].They madeavaluablepoint,however,thatsolidbehavioralevi- dencefortheexistenceofuniquehuesislacking.Huescaling resultswereputforwardbyseveralrespondentsasevidence supportingthedistinctionbetweenuniqueandintermediate hues[ 13 , 35 – 37 ].Resultsfromthepresentstudyundermine thisevidence.Broakes[ 38 ],alsoinresponsetoSaunders andvanBrakel,challengedcolorscientiststoperformexperi- mentsusinglime,purple,orange,andteal,andsuggestedthat “ ThefearsofSaundersandvanBrakelwillbejustifiedifpeo- plecanprovetodoaswellwithlime,purple,orange,andteal aswedowithred,yellow,green,andblue. ” Forhuescaling, surprisingly,thisturnsouttobethecase. Whatcanexplainthebehaviorofsubjectsinourhuescal- ingexperiment?Onepossibilityisthattheassumptionthat thereisconsensusforthesubjectiveuniquenessofred,green, blue,andyellowismisguided.Butanotherpossibility,which isimpossibletoruleoutinasubjectivetaskofthisnature,is thatsubjectsignoredormisinterpretedtheinstructionsand insteadperformedacolorsimilaritytask,ratingthesimilarity oftheprimaryandthetesthue.Butifthiscouldexplainthe resultsofthepresentstudy,itcouldalsoexplaintheformof allotherresultsfromhuescalingwhereuniquehuesaregiven asprimaries.Conclusionsfromhuescaling,oneofthemain linesofevidencethathasbeenadvancedinsupportofthe specialstatusofuniquehues,areunsound. B.Experiment2:EffectofInstructionsonSettingsof UniqueHues Wehavefoundthatsubstitutingintermediatecolortermsinto theinstructionsaltersmeansettingsofuniquehues.Ifsub- jectswereaskedforauniqueredthatwasneitherpurplish noryellowish,theyidentifiedamuchmoreorangeshadeof redthaniftheywereaskedforauniqueredthatwasneither bluishnoryellowish.Oursampleofnaïvesubjects,despitein mostcasesproducingcleanpsychometricfunctions[e.g., Fig. 5(b) ],didnotreliablyidentifyaparticularhueassubjec- tivelypure.Ourresultsallowforthepossibilitythatunique huesdonotexist,andthatsettingsofuniquehuesmightsim- plyreflectaperceptualintermediatebetweenthealternatives givenintheinstructions.However,wecannotexcludethe possibilitythatoursubjectsweresimplybiasedbythecon- tentsoftheinstructions.Whatwe can concludeisthatdecid- ingwhetheraparticularhueisneither “ toored ” nor “ too green ” tobeuniqueisnotasimpleenoughorsalientenough taskthatsubjectscanescapebias.Wecanalsoconcludethat thefactthatsubjectsareabletoidentifyaparticularblueas neithergreenishnorreddishisnotevidencethattheblueis unique. C.GeneralDiscussion Ourresultsundermineexistingbehavioralevidenceforthe specialstatusofuniquehues.Butourexperiments,andmost othersthathaveattemptedtoestablishtheuniquenessof uniquehuesempirically,employBrindleyclassBobserva- tions[ 39 ],wheresubjectsarerequiredtodescribethequality oftheirsensations.Onerecentandparticularlyinterestingex- ampleofaclassBmethodofinvestigatinguniquehuesis par- tialhuematching byLogvinenkoandBeattie[ 40 ],where subjectsarerequiredtoidentifythesetofhuesthatshares acomponentcolorqualitywithatesthue. Chromaticity classes (thelargestsetofstimulithatallpartiallymatch) areextractedfrommatricesofpartialhuematches,andfor about50%ofobserversfour “ unique ” huesthatbelongtoonly onechromaticityclassemerged.However,therewereseveral J.M.BostenandA.E.BoehmVol.31,No.4/April2014/J.Opt.Soc.Am.AA391 minorityphenotypes,includingsubjectswhoseemtohave morethanfouruniquehues,orwhohavenouniquehues. Thedifficultyofinterpretingthepossiblealternativestrategies ofsubjectswithatypicalresultshasledLogvinenko[ 41 ]to suggestthat “ thisproblemisfartooinvolvedtobelefttoinex- periencedobservers. ” However,itseemsequallylikelythat observerswhoareexperiencedinmakingcolorintrospec- tionsmightgiveinvalidorunrepresentativeresults.Even thosewhoarenotversedincolorsciencemayadoptanintro- spectivestrategyofbasingsubjectivejudgmentsonthe “ primary ” colorsthattheyareculturallyfamiliarwith. Giventhelimitationsofmethodsinvolvingintrospection, couldanypsychophysicalexperimenteitherproveordis- provetheexistenceofuniquehues?Whataboutperformance measuresorclassAobservations? Subjects ’ reliabilityatsettinguniquehuesisnoteasilycat- egorizedeitherasaclassBorasaclassAobservation.Inthe methodofaverageerror (whichconstitutesaclassAobser- vation),subjectsarerequiredtomakeamatchmanytimes, andthevarianceofmatchesistakenasameasureofperfor- mance.Measurementofthereliabilityofsettingsofunique huesusesthemethodofaverageerror,butthestandardto whichsubjectsaremakinga “ match ” isinternalratherthan external,sosubjectsarerequiredtojudgethequalityoftheir sensations.Ineithercase,itmightbethoughtreasonableto assumethatsubjectscansetuniquehuesmorereliablythan theycansetintermediatehues.However,Malkoc etal. [ 30 ] foundthattest – retestreliabilityisashighforbinary(inter- mediate)huesasitisforuniquehues.BostenandLawrance- Owen[ 29 ]haveindependentlyreplicatedthefindingsof Malkoc etal. [ 30 ]andhavefoundthatintra-subjectvariability (aswellasinter-subjectvariability)foridentifyinguniquehues isnodifferenttothatforidentifyingintermediatehues. AlthoughitisperhapstellingthatpureclassAobservations thatdistinguishtheuniquehueshave,untilrecently,been lacking(somestudieshavereportednegatively,thatunique huesarenotdistinguishedfromotherhuesusingclassAob- servations[ 29 , 42 – 46 ]),DanilovaandMollon[ 47 ]havefound thatthelocusofuniqueblueanduniqueyellowcorresponds tominimainthresholdsforcolordiscrimination.The correlationbetweenthelociofuniquehues,measuredusing subjectivejudgments,andapeakindiscriminationperfor- manceisinteresting.Suchfindingsdo,however,needtoac- countforthewiderangeofsettingsofuniquehuesfound acrossindividualsandacrossstudies[ 48 ].Acorrelationbe- tweenindividualdifferencesinthepositionsofcolordiscrimi- nationminimaandindividualdifferencesinsettingsofunique hueswouldbeprovocative. DanilovaandMollon ’ s[ 47 ]carefulmeasurements,and otherBrindleyclassAobservations,mightbeabletoestablish thepresenceorabsenceofchannelstunedtoparticularchro- maticities,buttheycannomoreestablishtheuniquenessof uniquehuesthanthemorewidespreadclassBobservations. Uniquenessisdefinedsubjectively,andthereforecannotbe proven.Perhapsitistimeforcolorscientiststoputunique huesaside,andknowthemsimplyasred,green,blue,and yellow. ACKNOWLEDGMENTS ThisworkwassupportedbyaResearchFellowshipfrom GonvilleandCaiusCollege,andbygrantEY011711 fromtheNationalInstitutesofHealthawardedtoDonald MacLeod. REFERENCES 1.E.Hering, ZurLehreVomLichtsinne (CarlGerold ’ sSohn, 1878). 2.L.M.HurvichandD.Jameson, “ Anopponent-processtheoryof colorvision, ” Psychol.Rev. 64 ,384 – 404(1957). 3.R.DeValois,I.Abramov,andG.Jacobs, “ Analysisof responsepatternsofLGNcells, ” J.Opt.Soc.Am. 56 ,966 – 977 (1966). 4.J.MollonandC.Cavonius, “ Thechromaticantagonismsof OpponentProcessTheoryarenotthesameasthoserevealed instudiesofdetectionanddiscrimination, ” in ColourVision DeficienciesVIII ,G.Verriest,ed.(Junk,1987),pp.473 – 483. 5.K.A.JamesonandR.G.D ’ Andrade, “ It ’ snotreallyred,green, yellow,blue:aninquiryintoperceptualcolorspace, ” in Color CategoriesinThoughtandLanguage ,C.HardinandL.Maffi, eds.(CambridgeUniversity,1997),pp.295 – 319. 6.I.AbramovandJ.Gordon, “ Colorappearance:onseeingred — or yellow,orgreen,orblue, ” Annu.Rev.Psychol. 45 ,451 – 485 (1994). 7.J.Mollon, “ Aneuralbasisforuniquehues? ” Curr.Biol. 19 , R441 – R442(2009). 8.C.M.StoughtonandB.R.Conway, “ Neuralbasisforunique hues, ” Curr.Biol. 18 ,R698 – R689(2008). 9.S.M.WuergerandL.Parkes, “ Uniquehues:perceptionandbrain imaging, ” in NewDirectionsinColourStudies ,C.Biggam,C. Hough,C.Kay,andD.Simmons,eds.(JohnBenjaminPublishing Company,2011),pp.445 – 455. 10.A.Valberg, “ Uniquehues:anoldproblemforanewgeneration., ” Vis.Res. 41 ,1645 – 1657(2001). 11.D.R.Hilbert, “ Basictastesanduniquehues, ” Behav.BrainSci. 31 ,82(2008). 12.M.DanilovaandJ.Mollon, “ Fovealcolorperception:minimal thresholdsataboundarybetweenperceptualcategories, ” Vis. Res. 62 ,162 – 172(2012). 13.D.Miller, “ Overtherainbow:theclassificationofuniquehues, ” Behav.BrainSci. 20 ,204 – 205(1997). 14.D.JamesonandL.M.Hurvich, “ Perceivedcoloranditsdepend- enceonfocal,surrounding,andprecedingstimulusvariables., ” J.Opt.Soc.Am. 49 ,890 – 898(1959). 15.C.E.SternheimandR.M.Boynton, “ Uniquenessofperceived huesinvestigatedwithacontinuousjudgmentaltechnique, ” J.Exp.Psychol. 72 ,770 – 776(1966). 16.R.L.DeValois,K.K.DeValois,E.Switkes,andL.Mahon, “ Hue scalingofisoluminantandcone-specificlights, ” Vis.Res. 37 , 885 – 897(1997). 17.V.J.Volbrecht,J.L.Nerger,L.S.Baker,A.R.Trujillo,andK. Youngpeter, “ Uniquehuelocidifferwithmethodology, ” OphthalmicPhysiol.Opt. 30 ,545 – 552(2010). 18.I.AbramovandJ.Gordon, “ Seeinguniquehues, ” J.Opt.Soc. Am.A 22 ,2143 – 2153(2005). 19.J.Gordon,I.Abramov,andH.Chan, “ Describingcolorappear- ance:hueandsaturationscaling, ” Percept.Psychophys. 56 , 27 – 41(1994). 20.B.WootenandD.Miller, “ Thepsychophysicsofcolor, ” in Color CategoriesinThoughtandLanguage ,C.HardinandL.Maffi, eds.(CambridgeUniversity,1997),pp.59 – 88. 21.M.A.Webster,E.Miyahara,G.Malkoc,andV.E.Raker, “ Variationsinnormalcolorvision.II.Uniquehues, ” J.Opt. Soc.Am.A 17 ,1545 – 1555(2000). 22.D.Hinks,L.M.Cárdenas,R.G.Kuehni,andR.Shamey, “ Unique- huestimulusselectionusingMunsellcolorchips, ” J.Opt.Soc. Am.A 24 ,3371 – 3378(2007). 23.G.JordanandJ.D.Mollon, “ Rayleighmatchesandunique green, ” Vis.Res. 35 ,613 – 620(1995). 24.F.L.DimmickandM.R.Hubbard, “ Thespectrallocationof psychologicallyuniqueyellow,greenandblue, ” Am.J.Psychol. 52 ,242 – 254(1939). 25.B.E.SchefrinandJ.S.Werner, “ Lociofspectraluniquehues throughoutthelifespan, ” J.Opt.Soc.Am.A 7 ,305 – 311(1990). 26.J.M.Eichengreen, “ Uniquehueloci:inducedshiftswith complementarysurrounds, ” Vis.Res. 16 ,199 – 203(1976). A392J.Opt.Soc.Am.A/Vol.31,No.4/April2014J.M.BostenandA.E.Boehm 27.D.H.Brainard,A.Roorda,Y.Yamauchi,J.B.Calderone,A. Metha,M.Neitz,J.Neitz,D.R.Williams,andG.H. Jacobs, “ Functionalconsequencesoftherelativenumbers ofLandMcones, ” J.Opt.Soc.Am.A 17 ,607 – 614(2000). 28.D.I.MacLeodandR.M.Boynton, “ Chromaticitydiagramshow- ingconeexcitationbystimuliofequalluminance, ” J.Opt.Soc. Am. 69 ,1183 – 1186(1979). 29.J.BostenandA.J.Lawrance-Owen, “ Nodifferenceinvariability ofuniquehueselectionsandbinaryhueselections, ” J.Opt.Soc. Am. 31 ,A357 – A364(2014). 30.G.Malkoc,P.Kay,andM.A.Webster, “ Variationsinnormal colorvision.IV.Binaryhuesandhuescaling, ” J.Opt.Soc. Am.A 22 ,2154 – 2168(2005). 31.K.ZychalukandD.H.Foster, “ Model-freeestimationofthepsy- chometricfunction, ” Atten.Percept.Psychophys. 71 ,1414 – 1425 (2009). 32.K.A.Jameson, “ WhereintheWorldColorSurveyisthesupport fortheHeringPrimariesasthebasisforColorCategorization? ” in ColorOntologyandColorScience ,J.CohenandM.Matthen, eds.(MIT,2010),pp.179 – 202. 33.P.KayandT.Regier, “ Resolvingthequestionofcolornaming universals, ” Proc.Natl.Acad.Sci.USA 100 ,9085 – 9089(2003). 34.B.A.SaundersandJ.vanBrakel, “ Aretherenontrivial constraintsoncolourcategorization? ” Behav.BrainSci. 20 , 167 – 179(1997). 35.I.AbramovandJ.Gordon, “ Constrainingcolorcategories:the problemofthebabyandthebathwater, ” Behav.BrainSci. 20 ,179 – 180(1997). 36.M.Bornstein, “ Selectivevision, ” Behav.BrainSci. 20 ,180 – 181 (1997). 37.A.ByrneandD.Hilbert, “ Uniquehues, ” Behav.BrainSci. 20 , 184 – 185(1997). 38.J.Broakes, “ Couldwetakelime,purple,orange,andtealas uniquehues? ” Behav.BrainSci. 20 ,183 – 184(1997). 39.G.Brindley, PhysiologyoftheRetinalandVisualPathway ,2nd ed.(CamelotLtd.,1970). 40.A.D.LogvinenkoandL.L.Beattie, “ Partialhue-matching, ” J.Vis. 11 (8):6(2011). 41.A.D.Logvinenko, “ Atheoryofuniquehuesandcolour categoriesinthehumancolourvision, ” ColorRes.Appl. 37 , 109 – 116(2012). 42.M.A.WebsterandJ.D.Mollon, “ Changesincolourappearance followingpost-receptoraladaptation, ” Nature 349 ,235 – 238 (1991). 43.C.WitzelandK.R.Gegenfurtner, “ Categoricalsensitivityto colordifferences, ” J.Vis. 13 (7):1(2013). 44.S.Burns,A.Elsner,J.Pokorny,andV.Smith, “ TheAbneyeffect: chromaticitycoordinatesofuniqueandotherconstanthues, ” Vis.Res. 24 ,479 – 489(1984). 45.G.MalkocandF.A.A.Kingdom, “ Dichopticdifferencethresh- oldsforchromaticstimuli, ” Vis.Res. 62 ,75 – 83(2012). 46.M.Ayama,T.Nakatsue,andP.K.Kaiser, “ Constanthuelociof uniqueandbinarybalancedhuesat10,100,and1000Td, ” J. Opt.Soc.Am.A 4 ,1136 – 1144(1987). 47.M.V.DanilovaandJ.D.Mollon, “ Cardinalaxesarenotindepen- dentincolordiscrimination, ” J.Opt.Soc.Am.A 29 ,A157 – A164 (2012). 48.R.Kuehni, “ Variabilityinuniquehueselection:asurprising phenomenon, ” ColorRes.Appl. 29 ,158 – 162(2004). J.M.BostenandA.E.BoehmVol.31,No.4/April2014/J.Opt.Soc.Am.AA393 Empiricalevidenceforuniquehues? J.M.Bosten 1,2, *andA.E.Boehm 1,3,4 1 DepartmentofPsychology,UniversityofCalifornia,SanDiego,McGillHall,LaJolla,California92093,USA 2 DepartmentofExperimentalPsychology,UniversityofCambridge,DowningStreet,CambridgeCB23EB,UK 3 VisionScienceGraduateGroup,UniversityofCalifornia,Berkeley,380MinorHall,Berkeley,California94720,USA 4 SchoolofOptometry,UniversityofCalifornia,Berkeley,380MinorHall,Berkeley,California94720,USA *Correspondingauthor:jmb97@cantab.net ReceivedOctober3,2013;revisedDecember22,2013;acceptedDecember27,2013; postedJanuary13,2014(Doc.ID198752);publishedMarch11,2014 Red,green,blue,yellow,andwhitehavebeendistinguishedfromotherhuesasunique.Wepresentresultsfrom twoexperimentsthatundermineexistingbehavioralevidencetoseparatetheuniquehuesfromothercolors.In Experiment1weusedhuescaling,whichhasoftenbeenusedtosupporttheexistenceofuniquehues,buthas neverbeenattemptedwithasetofnon-uniqueprimaries.Subjectswereassignedtooneoftwoexperimental conditions.Inthe “ unique ” condition,theyratedtheproportionsofred,yellow,blue,andgreenthattheyper- ceivedineachofaseriesofteststimuli.Inthe “ intermediate ” condition,theyratedtheproportionsofteal, purple,orange,andlime.Wefound,surprisingly,thatresultsfromthetwoconditionswerelargelyequivalent. InExperiment2,weinvestigatedtheeffectofinstructiononsubjects ’ settingsofuniquehues.Wefoundthat alteringthecolortermsgivenintheinstructionstoincludeintermediatehuesledtosignificantshiftsinthehue thatsubjectsidentifiedasunique.Theresultsofbothexperimentsquestionsubjects ’ abilitiestoidentifycertain huesasunique.©2014OpticalSocietyofAmerica OCIScodes: (330.1690)Color;(330.1720)Colorvision;(330.5510)Psychophysics;(330.5020)Perception psychology. http://dx.doi.org/10.1364/JOSAA.31.00A385 1.INTRODUCTION Fivehues:red,green,blue,yellow,andwhite,havebeengiven specialstatusbycolorscientistsas “ unique. ” Theuniquehues aresaidtobephenomenologicallypure,whileallotherhues aresaidtobephenomenologicallymixed,containingvarious proportionsofthefiveuniquehues. Uniquehueshavealonghistoryinpsychology.Itwasthe beliefthatuniquehuesmusthavespecialstatusincolorrep- resentationthatledHering[ 1 ]toproposered,green,blue,and yellowasthepolesofhisopponentmechanisms.Following thetraditionestablishedbyHering[ 1 ],andlaterbyHurvich andJameson[ 2 ],whencoloropponentcellswerediscovered intheprimatelateralgeniculatenucleus,theyweredescribed asbeingselectiveforredandgreen,andblueandyellow[ 3 ]. Althoughitisnowclearthatthechromatictuningofcolor- opponentmechanismsrevealedpsychophysicallydoesnot mapontotheHeringprimaries[ 4 ],andthatthecolor-sensitive cellsinthelateralgeniculatenucleusencodeintermediate colordirections[ 5 , 6 ],theconvictionthatuniquehuesmust haveprominenceinneuralcolorsignalshasledtopersistence oftheideathatcolor-opponentcellsencodetheHeringpri- maries,andtoasearchforhigher-ordercolormechanisms thatareselectiveforred,green,blue,andyellow[ 7 – 10 ]. A.HueScaling Despitebroadconsensusamongcolorscientiststhatred, green,blue,andyellowshouldbeconsideredunique,solid behavioralevidencetosupportthisviewhasbeenhardto comeby.Onesourceofevidencethatisoftencitedtojustify thespecialstatusofuniquehues[ 11 – 13 ],ishuescaling.Hue scalingasamethodisbasedonsensoryscalingofmultiple stimulusattributes,introducedbyJamesonandHurvich [ 14 ].Thesubjectispresentedwithcoloredstimuliandasked toconsiderhowmuchofvariousgiven “ primary ” colorseach stimuluscontains.Thesubjectassignsanumericalratingto eachprimarythatreflectshisjudgmentofhowmuchofthat primaryiscontainedinhisperceptionofthestimulusthatis presentedtohimonaparticulartrial.Theideaisthatifahue isunique,itwillcontainoneprimary,butnotanyother. Resultsfromhuescalingexperimentstypicallyshowthat therearefourpointsaroundthehuecirclewheresubjectsre- portseeingonlyoneprimary.Forexample,at “ unique ” green, subjectsreportseeinggreen,butnotblue,yellow,orred. However,sincetheprimariestowhichsubjectsareasked toassignratingsaretypicallythemselvestheuniquehues, itisperhapsunsurprisingthatratingspeakforred,green, blue,andyellowinregionswhereneighboringhuesapproach thezerolevel.Red,yellow,green,andbluearesetapart roughly90°aroundthehuecircle,soitmaybesimplytheper- ceptualdistancebetweenthemthatresultsinonenotbeing seeninanother. SternheimandBoynton[ 15 ]didhuescalingformonochro- maticlightsbetween530and620nm.Theyintroducedorange asanavailableprimaryinsomeoftheirconditions,andcon- cludedthattheyhadevidencetoseparateyellow,green,and red,butnotorange,asunique.Infourconditionssubjectswere given(i)red,green,andblue,(ii)red,yellow,andgreen,(iii)red, orange,andgreen,and(iv)red,orange,yellow,andgreen,touse asprimaries.Whenyellowwasunavailableasaprimary,there werecertainwavelengthsthatSternheimandBoynton ’ s[ 15 ] subjectswereunabletoadequatelydescribebymixturesof red,green,andblue.Incontrast,whenorangewasunavailable, J.M.BostenandA.E.BoehmVol.31,No.4/April2014/J.Opt.Soc.Am.AA385 1084-7529/14/04A385-09$15.00/0©2014OpticalSocietyofAmerica lightofwavelengthsinthe “ orange ” partofthespectrumcould beadequatelydescribedasmixturesofredandyellow. WebelievethatSternheimandBoynton ’ s[ 15 ]comparison betweenconditionswasnotentirelybalanced.Inconditions whereorangewasunavailableasaprimary,twocloselyneigh- boringhues,redandyellow,wereavailable.Butinconditions whereyellowwasmissing,atleastoneofthetwoalternative primariesprovidedwasfurtherfromyellowonthehuecircle thaneitherredoryellowfromorange.Itcouldsimplybethe spacingoftheprimariesincolorsimilarity,ratherthantheir uniqueness,thatallowsadequatedescriptionofotherhues. NeitherSternheimandBoynton[ 15 ]norotherswhohave sinceusedhuescalingtoinvestigatecolorappearance [ 16 – 20 ],haveintroducedatruecontrolcondition.Inorder toconcludethathuescalingresultsdemonstratetheunique- nessofuniquehues,itisnecessarytoshowthatthesamere- sultscannotbeobtainedusingonlynon-uniqueprimaries roughlyevenlydistributedaroundthehuecircle.InExperi- ment1,wereporttheresultsofhuescalingundertwodiffer- entconditions:astandard “ unique ” conditionwherethefour uniquehuesaregivenasprimaries,andacontrol “ intermedi- ate ” conditionwherefourintermediatehues(purple,orange, lime,andteal)aregiven. B.EffectofInstructiononSettingsofUniqueHues Inexperimentswheresubjectsarerequiredtoidentifyorpro- videsettingsofuniquehues,otheruniquehuesareinvariably givenintheinstructions[ 21 – 26 ].Asubjectmaybeaskedtoset ayellowthatisneitherreddishnorgreenish[ 27 ],orformea- surementsofuniquegreen,todecidewhetheragivencoloris toobluishortooyellowish[ 17 ]. Subjectsarerequiredtoidentifyauniquehuebyconsider- ationofother(typicallyunique)huesspecifiedintheinstruc- tions.Thetwootheruniquehuescontainedinstandard instructionsaresituatedroughlyorthogonallyaroundthe huecirclefromtheuniquehuetobeidentified.Thissuggests possibleartifacts.Redmaybeidentifiedasneitherbluishnor yellowishsimplybecauseblueandyellowareeachsuffi- cientlydistantfromred.Alternatively,redmaybeidentified asneitherbluenoryellowbecauseitisperceptuallyhalfway betweenblueandyellow,ratherthanbecauseitissubjectively pure.OurExperiment2controlsforthesepotentialexperi- mentalartifactsbysubstitutingnon-uniquecolortermsinto theinstructions.Insteadofbeingaskedtoidentifyaunique redthatisneitherbluishnoryellowish,subjectsmightbe askedtoidentifyauniqueredthatisneitherbluishnororang- ish.Ifsubjectsidentifyuniqueredasasinglesubjectivelypure color,thechangeininstructionshouldhavenoeffectontheir “ unique ” huesettings. 2.EXPERIMENT1:HUESCALING A.Predictions Ifuniquehuesaresubjectivelypureandotherhuesaresub- jectivelymixed,whatshouldtheresultsofhuescalingbefor ouruniqueandintermediateconditions?Predictionsare showninFig. 1 .Fortheuniqueprimariesweexpectfunctions likethosethathavebeenobtainedbymanyotherresearchers. Thefunctionforeachprimaryshouldpeakatthehueangle ofitsassociateduniquehue,wherethefunctionsforthe threeotherprimariesshouldbeatzero.Atthepositionsof intermediatehues,twoprimariesshouldbereported.The predictionfortheintermediateprimariesisverydifferent. Functionsforeachintermediateprimaryshouldpeakatthe hueangleofthatprimary.Butatthepositionsofthe(subjec- tivelyunitary)uniquehues,allfunctionsshouldbeatzero. B.Stimuli ArepresentationofaparticulartrialisgiveninFig. 2(a) .The teststimuluswasadiskof4°diameterpresentedintheupper halfofthescreen.Thechromaticityoftheteststimuluswas randomlyselectedoneachtrialfrom36chromaticitycoordi- natesdistributedinacircleinMacLeod – Boynton[ 28 ]chroma- ticityspacearoundapointmetamericwithilluminantD65 [Fig. 2(b) ].Intheresultssection,thechromaticitiesofthe 36stimuliwillbegivenby “ hueangles. ” Wedefinedhueangle inMacLeod – Boyntonchromaticityspaceclockwise,withzero atthevertical(S   L  M  )axis[Fig. 2(b) ].Conversionfrom hueangletoMacLeod – Boyntonchromaticitycoordinates canbeachievedbythefollowingformulas,where  isthe hueangle: S   L  M  0 . 0167  0 . 013 cos  ; L   L  M  0 . 6552  0 . 0364 sin  . Thebackgroundonwhichthestimuliwerepresentedwas metamericwithD65.Allteststimuliwereisoluminantwitha luminanceof 28 cdm  2 ,andtheluminanceofthebackground was 14 cdm  2 .Inthelowerpartofthescreen,disksof2° diameterwerepresentedthatcontainedfourprimaries [Fig. 2(b) ],whichwereeithertheuniquehuesortheinter- mediatehuesdependingonthecondition.Theluminance oftheprimarieswas 28 cdm  2 .Aboveeachprimarywasa boxinwhichthesubjectenteredanumbercorresponding tohisperceptionofhowmuchofthatprimarywascontained intheteststimulus.Theboxesweredelimitedwithablack line,onepixelwide.Thesubjectusedthearrowkeystomove betweenboxes,andthecurrentlyselectedboxwasindicated byreplacingtheblackborderwithawhiteborder,metameric Fig.1.Predictionsforhuescalingwithuniqueprimaries[panel(a)], andwithintermediateprimaries[panel(b)].Alongthe x axishuean- glesareplottedaroundthehuecircle,andonthe y axisthepredicted numericalratingsfrom0to9areshown.Thedashedverticallines indicatetheuniqueandbinaryhueprimaries,whicharelabeledwith uppercaseletters:purple(P),red(R),orange(O),yellow(Y),lime(L), green(G),teal(T),andblue(B).Predictedfunctionsforratingsfor eachprimaryasafunctionofthehueangleoftheteststimulus areindicatedbythesolidcurves,whicharelabeledwithlowercase letters. A386J.Opt.Soc.Am.A/Vol.31,No.4/April2014J.M.BostenandA.E.Boehm withD65andwithaluminanceof 28 cdm  2 .Subjectsentered aratingintoeachboxusinganumericalkeypad,andthe numberwasshownintheboxinwhite,alsometamericwith D65andwithaluminanceof 28 cdm  2 . C.SelectionofPrimaries Theprimariesusedinthehuescalingexperimentwerebased onselectionsofuniqueandintermediatehuesmadeby58sub- jects[ 29 ].Subjectswerepresentedwithannuliofselectable segmentscontainingaprogressionofhuesonaCRTmonitor coveredwithatouch-sensitivescreen.Thesegmentscontain- ingthehueswereisoluminantwithaluminanceof 28 cdm  2 . ThebackgroundwasmetamericwithD65andhadalumi- nanceof 17 cdm  2 .Oneachtrialacircleof90segmentscon- taining90discretehueswaspresented[Fig. 2(c) ].Thecircle wasrotatedrandomlyfromtrialtotrial.Accordingtothe block,subjectswereaskedtochoose,forexample, “ aredthat isneithertooorangenortoopurple ” or “ anorangethatisnei- thertoorednortooyellow ” [ 30 ].Thesubjectselectedthe segmenthethoughtbestmatchedtheinstruction,andasmall achromaticdiskappearedbesidethatsegment.Heconfirmed hisselectionbytappingachecksymbolpresentedinthelower leftpartofthescreen. Therewere16blocks,eachof15trials.Ineachblockoneof thefouruniquehues(red,green,blue,oryellow),oroneofthe fourintermediatehues(orange,purple,teal,orlime)was tested.Inthefirsteightblocks,alleighthuesweretested inarandomorder,andtheyweretestedagaininadifferent randomorderinthesecondeightblocks.Fifty-eightsubjects tookpartintheexperiment.Allhadnormalcolorvision assessedusingtheIshiharaplatespresentedunderMacBeth IlluminantC. Togatherresponsesofsubjects,weusedaKeytecMagic TouchProE-Xtouchscreen(modelnumberET2032C) attachedtoaCRTmonitor.Stimuliwerepresentedona DiamondPro2070SBCRTmonitor(Mitsubishi,Tokyo, Japan)calibratedusingaUDTphotometer(UnitedDetector Technology,Hawthorne,California)andaSpectraScan PR650spectroradiometer(PhotoResearchInc.,Chatsworth, California).ExperimentswereruninMatlab(TheMathWorks, Natick,Massachusetts)usingaVSG2/4graphicscard (CambridgeResearchSystems,Rochester,UK). Theprimariesusedinthehuescalingexperimentwere basedonthemeanhuesettingsforthe58subjects.These settingsareshowninFig. 2(d) alongwith95%confidence intervals. D.ProcedureforHueScaling Beforebeginningtheexperiment,subjectsweretoldthattheir taskwastomakesubjectivedecisionsaboutthecolorsofthe testdisks.Writteninstructionsfortheintermediatecondition wereasfollows: “ Youwillbeshownaseriesofcolors.For eachcoloryoumustdecidehowmuchoffourothercolors yoursubjectiveexperienceofthatcolorcontains.Youmust decidehowmuchorange,lime,purple,andtealyourexperi- enceofthecoloryouarepresentedwithcontains.Youmust ratethequantityofeachcolorinyourexperiencebyanumber from0to9.Enteryourratingsintotheboxesprovided.You cannavigatebetweenboxeswiththearrowkeys.Acceptall yourratingsforatrialbypressingReturn. ” Equivalentinstruc- tionsweregivenfortheuniquecondition,exceptthatthelist ofcolortermswas “ red,yellow,blue,andgreen. ” Subjectsweregivenadditionalinstructionsorally.They wereremindedthatthetaskwassubjective,thattherewas norightorwronganswerandthattheymustrespondinwhat- everwaytheyfeltwasmostappropriate.Theywereinstructed thattheratingstheyassigneddidnothavetoadduptoany particularnumberandthatanypossibleresponsewasaccept- able,includinggivingallfourprimariesascoreofzero. Oneachtrialasubjectenteredhisratingsintothefour boxes[Fig. 2(a) ].Hewasabletochangehisresponsesby overwritingthem.Whenhewassatisfiedwithallfourratings, hepressedtheReturnkeytomoveontothenexttrial.The36 testchromaticitieswerepresentedinarandomorderthree times,sothatall36teststimuliwerepresentedonceinaran- domorder,andthentwicemoreindifferentrandomorders. Fig.2.Stimuliusedinthehuescalingexperiment.Panel(a)repre- sentsthestimuluspresentedononetrial.Theupperdiskisthetest stimulus,andthefourlowerdisksaretheprimaries(lefttoright:red, yellow,green,andblue).Thesubjectwasinstructedtoassigna numericalratingtoeachprimaryaccordingtohowmuchofthatpri- maryheperceivedintheteststimulus.Theselectedboxwasindicated withawhiteborder.Thestimulusrepresentedherewasfortheunique condition:thestimulusfortheintermediateconditionwasequivalent, exceptthatthefourprimarieswereteal,lime,orange,andpurple. Panel(b)showsthechromaticitiesofthestimuliinMacLeod – Boynton chromaticityspace.Thetestchromaticitiesareshownbythedisks, andtheprimariesbythegraycrosses,labeled.Theblackdotin thecenterofthefigureindicatesthechromaticityofD65.Thehue angle(  )foranexampleredteststimulusisindicatedinthepanel. Panel(c)representsthestimulususedformeasuringuniqueandin- termediatehues.Eachof90segmentscontainedahuefromarange linearlyspacedaroundacirclecenteredatthecoordinatesofD65in MacLeod – Boyntonchromaticityspace.Instructionswerepresented intheupperpartofthescreen — fortheparticulartrialrepresented inthefigure,theinstructionsread “ pickanorangethatisneither toorednortooyellow. ” Thesubjectusedastylustoselectasegment, andanachromaticdiskappearedbesidetheselectedsegment.The selectionwasconfirmedbytappingthechecksign.Panel(d)shows meanselectionsofuniqueandintermediatehuesmadeby58subjects. Datapointsforeachprimaryarelabeled:purple(P),red(R),orange (O),yellow(Y),lime(L),green(G),teal(T),andblue(B).Errorbars (linesinsidedatapoints)indicate95%confidenceintervalsonthe mean.Themeanselectionswereusedastheprimariesinthehue scalingexperiment. J.M.BostenandA.E.BoehmVol.31,No.4/April2014/J.Opt.Soc.Am.AA387 Asubject ’ sratingforeachteststimuluswasbasedonthe meanofthethreetrials.Theexperimenttookmostsubjects between10and30minutestocomplete. E.Subjects Thirty-sixsubjectstookpartintheexperiment.Allhadnormal colorvisionassessedusingtheIshiharaplates.Subjectswere assignedrandomlyeithertothe “ unique ” ortothe “ intermedi- ate ” conditionsothat18subjectsparticipatedineachcondi- tion.ThesubjectswereUniversityofCalifornia,SanDiego undergraduatestudentswhoparticipatedinexchangefor coursecredit.Theystudiedarangeofsubjects,including statistics,psychology,biology,andlinguistics.Allsubjects werenaïvetothepurposesofthehuescalingexperiment andhadnottakenpartintheexperimenttoselectthe chromaticitiesoftheprimaries. F.Equipment StimuliwerepresentedonaMitsubishiDiamondPro2070SB CRTmonitor.Thegammafunctionswerelinearizedusing measurementsmadewithaUDTphotometercalibratedfor luminanceusingaSpectrascanPR560spectroradiometer. Thecolorcalibration,whichallowedforconversionfrom MacLeod – BoyntonchromaticitycoordinatestoRGBvalues, wasachievedbymeasuringthespectraofthethreephosphors oftheCRTmonitorusingtheSpectrascanPR650. G.Results Resultsaveragedacrosssubjectsarepresentedin Figs. 3(a) – 3(b) .Thefigureshowsthemeanratingsforhow muchofeachprimarywasjudgedtobecontainedineachtest stimulus.Resultsfromtheuniqueconditionarepresentedin panel(a),andresultsfromtheintermediateconditionarepre- sentedinpanel(b).Inbothpanels,thehueanglesoftheeight primariesusedinthetwoconditionsareindicatedbythever- ticaldashedlines. Resultsfortheuniqueconditionweresimilartothoseob- tainedbySternheimandBoynton[ 15 ].AsSternheimand Boynton[ 15 ]found,peakratingsareatthehueanglesof theuniquehues.Attheangleofeachuniquehue,ratings forthethreeotheruniquehueswereverylow.Attheangles ofintermediateprimaries,twouniqueprimariesweregiven mediumratings. Surprisingly,theresultsoftheintermediateconditionwere verysimilartothoseoftheuniquecondition.Peakratingsfor eachintermediateprimarywerefoundatthehueangleofthat primary.Attheangleofeachintermediateprimary,ratingsfor thethreeotherintermediateprimarieswerelow.Attheangles Fig.3.Resultsofthehuescalingexperiment.Ineachpanel,thehueangles(indegrees)oftheteststimulishowninFig. 2(b) areplottedalongthe x axesandratingsareplottedupthe y axes.Eachfunctionshowsratingsforhowmuchofaparticularprimarysubjectsjudgedtobepresentinthe teststimuli.Eachpanelshowsfourfunctionsdescribingtheresultsforthefourprimariesprovidedinthatcondition,coloredaccordingtothe primary.Dashedverticallinesindicatethehueanglesoftheeightprimariesusedacrossthetwoconditions,alsocoloredaccordingtotheprimary. Panel(a)showsgroupmeanresultsfortheuniquecondition( n  18 ),andpanel(b)showsgroupmeanresultsfromtheintermediatecondition ( n  18 ).Errorbarsinpanels(a)and(b)are95%confidenceintervalsforeachmeanrating.Panels(c) – (h)showexamplesofresultsfromindi- vidualsubjects,withthengivenforeachcaseindicatingthenumberofsubjectswhoproducedaresultsimilartotheoneshown.Panels(c) – (e)are fromtheuniquecondition,andpanels(f) – (h)arefromtheintermediatecondition. A388J.Opt.Soc.Am.A/Vol.31,No.4/April2014J.M.BostenandA.E.Boehm oftheuniqueprimaries,twointermediateprimarieswere givenmediumratings. Therewereothernotablefeaturesofthedata.Ineachcon- dition,theintersectionoftwodescendingfunctionsoccurred verynearthehueangleofaprimarynotusedinthatcondition. Forexample,intheuniquecondition,thefunctionsforthered andyellowprimariescrossedatabout125°,whichwas veryclosetothehueangleoftheorangeprimarythatwas notprovidedintheuniquecondition.Intheintermediate condition,thefunctionsforthelimeandorangeprimaries crossedatabout160°,whichwasclosetothehueangle fortheyellowprimarythatwasnotprovidedintheintermedi- atecondition. Themoststrikingfeatureoftheresultsisthesimilaritybe- tweenthetwoconditions.Differencesbetweenconditions,by comparison,arerelativelyminor.However,the “ baseline ” rat- ings(ratingsforprimariesatteststimuluspositions180°re- movedfromthepositionwhereratingspeak)arealittlehigher intheintermediateconditionthanintheuniquecondition. Thissmalldifferencebetweenconditionsmaybedrivenby individualdifferences.Aminorityofsubjectsperformed unusuallyonthistask,anditmaybethatinthetwoconditions thereweredifferentnumbersofsubjectswhofollowedminor- itystrategies. H.IndividualDifferences Resultsfromsixindividualsubjectsareshownin Figs. 3(c) – 3(h) .Resultsoftheuniqueconditionarepre- sentedinpanels(c) – (e),andresultsoftheintermediatecon- ditionarepresentedinpanels(f) – (h).Ineachpanel,n indicatesthenumberofsubjectswhoproducedresultssim- ilartotheexampleshown.Mostsubjectsassignedhighrat- ingstooneortwoprimariesforeachteststimulus,andzero ornearzeroratingstotheotherprimaries[panels(c)and (f)].Othersubjects,tovaryingdegrees,assignednon-zero ratingstomorethantwoprimariesforagiventeststimulus [panels(d)and(g)].Twosubjects[panels(e)and(h)]gave resultsthatresembledthepredictionsforintermediatehues [Fig. 1(b) ],butoneofthesewasgivenuniqueprimaries. Theaveragevariabilityinaratingwasgreaterforthe intermediatethanfortheuniquecondition.Specifically, themeanstandarddeviationofratingswas1.59forthein- termediateconditionand1.29fortheuniquecondition ( T  5 . 9 , p 0 . 01 ).However,ratingvariabilityonlydiffered betweenconditionsforlowmeanratings(Fig. 4 ).At mediumandhighmeanratings,thedistributionsofvariabil- ityinratingsoverlapped.Thisshowsthatthegreatervari- abilityintheintermediateconditionwasonlynearthe “ baseline ” levelofrating.Thisgreatervariationinbaseline ratingsindicatesthattheslightlyhigherbaselineratingsin theintermediateconditionarecausedbyindividual differences.Figures 3(c) – 3(h) bearthisout:moresubjects assignednon-zeroratingstomorethantwoprimariesfor agiventeststimulusintheintermediateconditionthan intheuniquecondition. Thesmalldifferenceinresultsbetweenthetwoconditions appearstobedrivenbyindividualdifferencesintaskstrategy. Therewereclearexamplesofthreedifferentstrategiesinboth conditions[Figs. 3(c) – 3(h) ],soweconcludethatprovidingin- termediateratherthanuniqueprimariesforhuescalingcan leadtoequivalentresults. 3.EXPERIMENT2:EFFECTOF INSTRUCTIONONSETTINGSOFUNIQUE HUES A.Stimuli Thestimuluswasadiskofapproximately2°presentedinthe centerofthedisplay.Theluminanceofthediskwas 28 cdm  2 .Itschromaticityvariedaccordingtoastaircasepro- ceduretrackingthesubject ’ sresponses.Thestartingpositions ofthestaircaseswereathueanglesof0°(L   L  M  0 . 655 , S   L  M  0 . 0297 ),and270°(L   L  M  0 . 619 ,S   L  M  0 . 0167 )whenuniquebluewasmeasured;andat 0°(L   L  M  0 . 655 ,S   L  M  0 . 0297 )and140° (L   L  M  0 . 679 ,S   L  M  0 . 00675 )whenuniquered wasmeasured.ThebackgroundwasmetamericwithD65 andhadaluminanceof 21 cdm  2 .Stimuliwerepresented onthesamecalibratedMitsubishiDiamondProCRTmonitor asinExperiment1. B.Procedure Subjectswereassignedrandomlytooneofthreeconditions foruniqueblueandtooneofthreeconditionsforuniquered. Subjectswereinstructedthattheirtaskwastomakesubjec- tivedecisionsaboutthecolorofthetestdisk.Foruniqueblue theyweretheninstructed(dependingontheconditionto whichtheywereassigned)asfollows: “ Inthisblockyou arelookingforapurecolorthatis ” b(i) “ neithergreenishnorreddish, ” b(ii) “ neithertealishnorreddish, ” b(iii) “ neithergreenishnorpurplish. ” Theywereinstructedtopressonebuttononakeypadif,for example,thediskwas “ toogreen, ” andanotherbuttonifit was,forexample, “ toopurple. ” Foruniquered,subjectswere givenequivalentinstructions,exceptthecolorwordsgivenin thethreeconditionswere: r(i)blueandyellow, r(ii)blueandorange, r(iii)purpleandyellow. Bothuniqueblueanduniqueredweremeasuredforeach subjectinseparateblocksinarandomorder. 0 2 4 6 8 10 0 0.5 1 1.5 2 2.5 3 Mean rating Standard deviation of ratings Fig.4.Distributionsofratingvariabilitywithmeanrating.Standard deviationsofratingsfortheintermediateconditionareshownbythe graysquaresandfortheuniqueconditionbytheblackcircles.The distributionsofvariabilityinratingslargelyoverlap.However,lowrat- ingsshowlowervariabilityintheuniqueconditionthanintheinter- mediatecondition. J.M.BostenandA.E.BoehmVol.31,No.4/April2014/J.Opt.Soc.Am.AA389 Ineachblock,fourrandomlyinterleavedstaircases convergedonasubject ’ suniquehue.Theinitialstepsize wasahueangleof10°,andthiswasreducedto5°oncea pairofstaircaseshadcrossed[seeFig. 5(a) ].Eachstair- caseterminatedafter60trials.Therewere240trialsper block. C.Subjects Onehundredfifteensubjectstookpartintheexperiment. NonehadtakenpartinExperiment1.Allhadnormalcolor visionassessedusingtheIshiharaplates.Allsubjectswere naïvetothepurposesoftheexperiment.TheywereUniversity ofCalifornia,SanDiegoundergraduateswhoparticipatedin exchangeforcoursecredit. D.Results ResultsofExperiment2areshowninFig. 5 .Panel(a)shows examplesofstaircasesforuniqueblue.Thesubjectwhose dataareshowninthepanelwasgiveninstructionsb(ii),to identify “ abluethatisneithertealishnorreddish. ” Panel (b)showsapsychometricfunctionfittothedatashownin panel(a).WefitcumulativeWeibullpsychometricfunctions using modelfreev1.1 [ 31 ].Uniquehuesettingsweredefinedas the50%pointonthepsychometricfunction,wheresubjects wereequallylikelytogiveeachofthetwoalternativere- sponses.Panel(c)showsgroupresultsplottedinthecontext ofthefullhuecircle.Whitelinesshowgroupmeanhue settings,andthelightgrayareasindicate95%confidence intervals.Hueanglesofthehuesettingsareshowninpanel (d)foruniqueredandpanel(e)foruniqueblue. Thedatashowthatmeansettingsofuniquehuesareshifted bychangesintheinstructionsthatweregiventosubjects. Therewasasignificanteffectofinstructionbothforunique red( F  22 . 7 , p  5 . 6 × 10  9 )andforuniqueblue( F  64 , p 1 × 10  16 ).Inbothcases,huesettingsforthestandardin- structionswereintermediatebetweenthosefortheothertwo conditions.Comparedtosettingsresultingfromthestandard instructions, “ abluethatisneithergreenishnorreddish, ” set- tingsof “ abluethatisneithergreenishnorpurplish ” were shiftedawayfrompurpleandsettingsof “ abluethatisneither tealishnorreddish ” wereshiftedawayfromteal.Thepattern ofresultsforuniqueredwasanalogoustothatforuniqueblue. 4.DISCUSSION A.Experiment1:HueScaling Wehavefoundthat,surprisingly,whenasetofintermediate huesaresubstitutedfortheuniquehuesasprimariesinhue scaling,resultsremainbroadlythesame.Ourresultsare clearlyagainstpredictionsofwhatshouldhappengiventhe prevailingviewthatuniquehuescanbedistinguishedfrom otherhuesassubjectivelypure(compareFigs. 1 and 3 ).Sub- jectsreportedseeingintermediateprimariesin “ unique ” hues inthesamewayastheyreportedseeinguniqueprimariesin intermediatehues.Intheintermediatecondition,subjectsas- signedratingsofaboutfivetoeachofthepurpleandorange Fig.5.ResultsofExperiment2.(a)Examplesofstaircasesforonesubjectinconditionb(ii): “ Neithertealishnorreddish. ” (b)CumulativeWeibull psychometricfunctionfittothedatashownin(a).Theareaofeachdatapointisproportionaltothenumberoftimestheparticularstimuluswas presented.(c)Groupresults.Whitebarsindicategroupmeansforeachcondition,andlightgrayareasindicate95%confidenceintervals.Ineach case,the ‘ standard ’ versionofthequestion(conditioni)producesintermediateresults,withalterationsinthecolorwordsgivenintheinstructions predictablyshiftingthemeansettingsofuniquehues(conditionsiiandiii).(d)Resultsforuniqueredshownasafunctionofhueangle.Errorbars are95%confidenceintervals.(e)Resultsforuniqueblueshownasafunctionofhueangle.Errorbarsare95%confidenceintervals. A390J.Opt.Soc.Am.A/Vol.31,No.4/April2014J.M.BostenandA.E.Boehm