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Crustaceana866718727


HYASTENUSBARUANEWSPECIESOFSPIDERCRABFROMINDONESIABRACHYURAMAJOIDEAEPIALTIDAEWITHAKEYTOTHESPECIESOFHYASTENUSAMANDAMWINDSORandSHANETAHYONGAustralianMuseum6CollegeStreetSydneyNSW2010AustraliaABSTRACTAnew

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1 Crustaceana86(6)718-727 HYASTENUSBARU,AN
Crustaceana86(6)718-727 HYASTENUSBARU,ANEWSPECIESOFSPIDERCRABFROMINDONESIA(BRACHYURA,MAJOIDEA,EPIALTIDAE)WITHAKEYTOTHESPECIESOFHYASTENUSAMANDAM.WINDSORandSHANET.AHYONGAustralianMuseum,6CollegeStreet,Sydney,NSW2010,AustraliaABSTRACTAnewspeciesofspidercrab,Hyastenusbarun.sp.,wascollectedfromdeepwaterofftheeasternIndonesianislandKei-BesarbytheFrench-IndonesianKARUBARexpeditionin1991.Thenewspeciesissimilartoseveralothermembersofthegenus,butdiffersinspinationofthecarapaceandambulatorypereopods.Unenouvelleespècedecrabearaignée,Hyastenusbarun.sp.,aétécollectéeeneauprofondeaulargedel’îleest-indonésiennedeKeiBesar,parl’expéditionfranco-indonésienneKARUBARen1991.Cettenouvelleespèceressembleàplusieursautresespècesdugenre,maisendiffèreparlarépartitionetlenombred’épinesdelacarapaceetparlespéréiopodesambulatoires.INTRODUCTIONTheepialtidspidercrabgenusWhite,1847currentlyincludes38validspecies(Ngetal.,2008)thatweredividedintothreegroups,namedGroups1-3basedongonopod1morphologybyGrifn&Tranter(1986).MostspeciesbelongtoGroup1,recognisedbytherelativelystraightgonopod1withasubterminalapertureandconvexlobebehindtheapex.AnundescribedGroup1specieswascollectedfromtheBandaSeaneartheKeiIslands,MalukuProvince,IndonesiaaspartoftheKARUBARexpeditiontoIndonesiain1991.Thisspecies,knownfromasinglemalespecimen,issufcientlydistinctfromotherspeciesoftowarrantitsrecognitionasanewspecies,describedbelow. Correspondingauthor;e-mail:amwindsor@gmail.comKoninklijkeBrillNV,Leiden,2013DOI:10.1163/15685403-00003202 HYASTENUSBARUNOV.MATERIALANDMETHODSMorphologicalterminologygenerallyfollowsGrifn&Tranter(1986).Cara-pacelength(cl)ismeasureddorsallyfromtherostralapicestotheposteriormarginofthecarapace,excludingtheintestinalspine.Postrostralcarapacelength(pcl)ismeasuredalongthedorsalmidlinefromthebaseoftherostralsinustotheposteriormarginofthecarapace,excludingtheintestinalspine.Carapacewidth(cw)isthegreatestcarapacewidthandexcludestheepibranchialspines.Thespecimenisde-positedintheMuséumnationald’Histoirenaturelle,Paris(MNHN).ComparativematerialofselectedspeciesofspeciesintheAustralianMuseum(AM)isgivenintheAppendix.F

2 orfuturecomparativepurposes,thesequenceo
orfuturecomparativepurposes,thesequenceoftheCOIDNAbarcoderegionhasbeendepositedinGenBankunderaccessionnumberTAXONOMYPIALTIDAEMacLeay,1838ISINAEDana,1851Hyastenusbarun.sp.(gs.1-2)Materialexamined.—Holotype:MNHN-IU-2012-994,male(cl11.5mm,pcl7.8mm,cw4.7mm),BandaSea,KeiIslands,westofKei-Besar,KARUBARstnDW22,5°22S133°0185-124m,25October1991.Diagnosis.—Carapacepyriform,gastricandcardiacregionsstronglyelevated,bothlackingapicaltubercle;ablunt,medialtubercleonanteriorfaceofgastricelevation;low,indistinctlateralprotogastrictubercle;strong,lateralbranchialspineand2smalldorsaltuberclesinlongitudinalrow;pterygostomianmarginwithrowof2tuberclesfollowedby1sub-branchialtubercle.Rostralhornsdivergentforentirelength,hornshalfaslongaspostrostralcarapacelength.Anterolateralspineoffusedbasalantennalarticlewithmedian,dorsallydirectedaccessoryspinulevisibleindorsalview.Pereopod2-4meriwith2or3spinesonproximalhalfofextensormargin.Description.—Carapacepyriform,1.6timesaslongaswide,regionsdened;sparselycoveredwithshorthookedsetae.Rostralhornsdivergentforentirelength,horizontalinlateralview,lengthhalfpostrostralcarapacelength;marginslinedwithshort,soft,lobularsetaeandscatteredsimplesetae.Orbitaleaveweaklyexpanded,preorbitalangleproducedasastout,anterodor-sallydirectedspine;antorbitalangleproducedasaveryshort,bluntlyangularlobe;postorbitallobeshort,anteriorlycupped,laterallytruncate;dorsalorbitalhiatusa AMANDAM.WINDSOR&SHANET.AHYONGshallow,narrowU-shape;orbitopenventrally,eyestalkunprotected;hiatusbroad,Gastricregionelevated,withoutapicaltubercle;alow,blunttubercleanteriortopeakofgastricelevation;protogastricregionwithlowindistincttuberclelaterally;urogastricregionat,lackingtubercles.Hepaticregionslightlyinated,withsmall,lowtubercleonlateralmargin.Pterygostomianregionwith2tuberclesinlongitudinalrowbelowmoultsuture.Branchialregionslightlyinated;lateralmarginwithstraight,laterallydirectedepibranchialspineatlevelofpereopod3;dorsalsurfacewith2short,conicaltuberclesinlongitudinalrow,anteriortubercleatlevelofpereopod2,posteriortubercleatlevelofpereopod3,mesialtolateralepibranchialspine.Branchials

3 ub-marginwith1tubercleanteriorly,slightl
ub-marginwith1tubercleanteriorly,slightlybelowmoultsuture,inlinewithpterygostomiantubercles.Cardiacregionelevated,smooth,conical,unarmed,apexbluntlyrounded,slightlyhigherthangastricregion.Intestinalregionwithshort,conicalposterodor-sallydirectedspine,withshortsetae.Eyestalksshort,sparselysetoseanteriorly,corneaterminal.Basalantennalarticleabouttwiceaslongaswide;anterolateralspinepromi-nent,visibleindorsalview;lateralmarginwithshortanterodorsallydirectedspin-ule;lateralmarginconvex,slightlyexpandedundereyestalk;prominenttuberclesituatedbetweenantennalglandandmarginofventralorbitalhiatus.Penultimateantennalarticleoverreachingbaseofrostralsinusbydistalhalf;ultimatearticleabout0.6lengthofpenultimatearticle;marginsofbotharticleswithsmall,scat-teredbulboussetae.Maxilliped3smooth,merustriangular,anterolateralanglestronglyproduced,apexrounded,broaderthanischium;ischiumsubquadrate,mesialmargin12-dentate,outersurfacewithshallowlongitudinaldepression.Cheliped(pereopod1)slender,about0.9pcl,withscatteredbulbousorclub-shapedsetae;merusslightlyshorterthanpropodus,extensormarginwith2proximalspinesandstrongdistalspine;carpusaboutaslongasdactylus,withsmalltubercleatdistalthirdoflateralmargin;propodussmooth,dactylusandpollexequal,shorterthanone-halfpalmlength,occlusalmarginscrenulate,proximalgapeslight.Ambulatorylegsslender;pereopod22.0pcl;pereopod31.2pcl;pereopod4aslongaspcl;pereopod50.9pcl;sparselycoveredwithshortsimplesetaeandshort,club-likesetae.Pereopod2and3merusextensormarginwithprominentdistalspineand3smallerproximalspines;dactylusexormarginwith10or11(pereopod2)or7(pereopod3)well-spacedspines,distal2perpendiculartodacty-laraxis,remainderslightlyinclinedproximally.Pereopod4merusextensormarginwithshortdistalspineand2smallerproximalspines;dactylusexormarginwith HYASTENUSBARUNOV.7well-spacedspines,distal2perpendiculartodactylaraxis,remainderslightlyin-clinedproximally.Pereopod5merusextensormarginwithbluntdistalangleand1proximalspine;dactylusexormarginwith6well-spacedspines,perpendiculartodactylaraxis.Maleabdomenwith6freesomitesandtelson,widestatsomites2and3;surfacesparselycovered

4 withshortbulboussetae;segment1withamedia
withshortbulboussetae;segment1withamedialelevation,somite3withroundedelevationsnearlateralmargins.Telsontriangular,aslongaswide,marginsstraight,apexrounded.Gonopod1(malepleopod1)straight,attened,taperingtoacutedistalpoint;aperturesubdistal,lateralmarginwithweaklybilobedsubdistalswelling.Etymology.—Thespeciesnamemeans“new”inBahasaIndonesia,acknowl-edgingthatthisisanewspeciesfromIndonesia.Thenameisusedhereasifitwereanouninapposition.Remarks.—OfthespeciesinGroup1(Grifn&Tranter,1986),HyastenusbarumostcloselyresemblesH.ambonensisGrifn&Tranter,1986,H.brockiiDeMan,1887,H.subinermisZehntner,1894,andH.uncifer1909onthebasisofthesizeandpositionofbranchialtuberclesandspinesandtherelativelengthanddivergenceoftherostralhorns.HyastenusbaruH.ambonensis(AMP34609;Grifn&Tranter,1986:g.39)bothhaveelevatedcardiacandgastricregions,butthecardiacregionislowerthanthegastricregioninH.ambonensis.Bothspeciesalsosharethepresenceofamediantubercleontheanteriorsurfaceofthegastricelevation,butthistubercleisprominentinH.ambonensisandshortandbluntinH.baru(g.1B).ThelateralprotogastrictuberclesareconspicuousinH.ambonensis,asarethetuberclesattheapicesofthegastricandcardiacelevationsbuttheseareobscureandabsentinH.baru,respectively.Thedistalangleoftheambulatorypereopodmeriisabluntprojection,notasharpspineasinH.baru(g.2F).TherearenoproximalmeralspinesonanypereopodinH.ambonensis(cf.Grifn&Tranter,1986,g.39e).Hyastenusambonensisoverlapsindepthrangewith,withthetypeofH.ambonensishavingbeencollectedfrom90m(Grifn&Tranter,1986).ThecardiacandgastricregionsinH.brockii(AMP34616,P34617,P34618;Grifn&Tranter,1986,g.38h)arelow,withthecardiacregionnothigherthanthegastricregion,asopposedtothestrongelevationoftheseregionsinH.baruThereisnomedialtubercleontheanteriorsurfaceofthegastricelevationinbrockii,butthelateralgastrictuberclesarenotobscuredasinH.baru(g.1B).InbothH.brockiiH.baru(gs.1A,2A,B)thereisasmallaccessoryspinuleontheanterolateralspineofthebasalantennalarticle,bothofwhicharevisibleindorsalviewinbothspecies.InH.brockii,thereisaterminalspineonpereopod2andasmallproximalspineontheexormarginofthesamepereop

5 od,butnoton AMANDAM.WINDSOR&SHANET.AHYON
od,butnoton AMANDAM.WINDSOR&SHANET.AHYONG Fig.1.Hyastenusbarusp.nov.,maleholotype,cl11.5mm,cw4.7mm(MNHN).A,dorsalhabitus;B,carapace,rightlateralview.Scalebar1.0mm. HYASTENUSBARUNOV. Fig.2.Hyastenusbarusp.nov.,maleholotype,cl11.5mm,cw4.7mm(MNHN).A,rightantennalandorbitalregion,dorsalview;B,leftantennalandorbitalregion,ventralview;C,leftmaxilliped3;D,abdomen;E,rightpereopod1(cheliped);F,rightpereopod2merus;G,rightpereopod2dactylus;H,rightpereopod3dactylus;I,rightpereopod4dactylus;J,rightpereopod5;K,rightgonopod1,abdominalview;L,rightgonopod1,sternalview;M,tipofrightgonopod1,lateralview;N,tipofrightgonopod1,sternolateralview.Scalebar:A-J2.0mm,K-N0.2mm. AMANDAM.WINDSOR&SHANET.AHYONGanyotherpereopod.Hyastenusbrockiiisreportedfromdepthsof8-54m(Grifn&Tranter,1986).HyastenusbarushareswithH.subinermis(AMP34639;Loh&Ng,1999:gs.6-8)arostrumthatisapproximatelyhalfpcl,noapicaltuberclesoneitherthegastricorcardiacelevations,andobscurelateralgastrictubercles.Thespeciesdifferintheabsenceofamedialgastrictubercle,andtheabsenceofbothproximalanddistalspinesonallambulatorypereopodsinH.subinermis.InH.subinermisthegastricandcardiacregionsarebothlowandnotconicallyelevatedasinHyastenussubinermis,whichwasshowntobeaseniorsynonymofRathbun,1916byLoh&Ng(1999),isreportedfromdepthsof18-34m(Grifn&Tranter,1986;Rathbun,1916).TherostrumofH.uncifer(AMP34640;Grifn&Tranter,1986,gs.38e,f,i,39c,42e-g)isequaltothepclcomparedtoone-halfpclinH.baru.Whereasdoesnothavestronglyelevatedcardiacorgastricregions,theseregionsdohaveanapicaltubercleincontrasttothestrongelevationandabsenceofapicaltuberclesinH.baru.ThelateralgastrictuberclesaremoreprominentinH.unciferthaninH.baru(g.1A).Hyastenusunciferisreportedfromshallowerdepths(13-15m)(Grifn&Tranter,1986)thanH.baruAkeytothespeciesofisgivenbelow,modiedfromGrifn&Tranter(1986).Group1speciesaremarkedinboldface.KEYTOTHESPECIESOFTHEGENUSHYASTENUS1.Carapacestronglytuberculate;protogastricandmesogastricregionswith13ormoretubercles;cardiacregionwith6ormoretubercles....................................2–Carapacemoderatelyorweaklytuberculateorsmooth;protogastricandmesogastri

6 cregionswith0-7tubercles;cardiacregionwi
cregionswith0-7tubercles;cardiacregionwith0-5tubercles..................................32.Preorbitalangleofeavenotproduced,rounded;anterolateralangleofbasalantennalarticlenotproduced.Tuberclesondorsalpartofhepaticregionsubequaltodorsalbranchialtubercles;lateralbranchialtuberclesub-equaltobranchialtuberclemedialtoit...........H.whiteiGrifn,1976–Preorbitalangleofeavesharp,weaklyproducedorsometimesashortspine;anterolateralangleofbasalantennalarticleproducedasabluntlobeorspine.Tuberclesondorsalpartofhepaticregionsmallerthandorsalbranchialtubercles;lateralbranchialtubercleorspinehigherthanbranchialtuberclemedialtoit.........................H.sebaeWhite,18473.Upperorbitalhiatusverynarrow,U-shaped(almostslit-like),shallow;lowerorbitalhiatusashallowslit......................................................................4–UpperorbitalhiatusU-shapedorkeyhole-shaped,notparticularlynarroworshallow;lowerorbitalhiatusbroadlyU-shaped.....................................................64.Mesogastricregionwithalargetubercleatthesummitandaspineorlargetubercleontheanteriorslope...............................................H.aries(Latreille,1825)–Mesogastricregionwithatubercleatthesummit,theanteriorslopesmoothorwithalowtubercle..........................................................................5 HYASTENUSBARUNOV.5.Branchialregionwithashortlateralbranchialspine(0.1cw)andatuberclemedialandanteriortoit................................................H.auctusRathbun,1916–Branchialregionwitharobustlateralbranchialspine(1/6to1/4cw)butlackingatuberclemedialtothespine...................................H.elatusGrifn&Tranter,19866.Lateralmarginofbasalantennalarticlewithtwolobesofsimilarshapeandsize,sometimes,butnotalwaysparallel.............................................................7–Lateralmarginofbasalantennalarticlewithanterolaterallobelownarrow(orlacking)andmarginbehinditwithabroad,weaktostrong,convexityorlobeorelsenearlystraight..107.Rostralspinesfusedforbasalthird.Intestinalregionwithastrong,bluntposteriorspine...H.mindoroGrifn&Tranter,1986–Rostralspinesseparatefromthebas

7 e.Intestinalregionwitharoundedtubercle,s
e.Intestinalregionwitharoundedtubercle,sometimesverylow..........................................................................88.Rostralspinesveryshort,about1/6pcl.Dorsalbranchialmarginwith5-9tubercles,epibranchialtuberclehardlyhigherthanothertubercles......H.fraterculusRathbun,1916–Rostralspines1/4to1/2pcl.Dorsalbranchialmarginwith2-4tuberclesandanepibranchialspinehigherthanthetubercles......................................................99.Cardiacregionwithapairofsubmedialtubercles.LobesonlateralmarginofbasalantennalarticleapicallyroundedH.biformisRathbun,1916–Cardiacregionwithamedialtubercle.Lobesonlateralmarginofbasalantennalarticlesubacute..............................................H.scrobiculatusRathbun,191610.Anteriorhalfofbranchialregionwithatleast1dorsaltubercle(sometimeslow)........11–Anteriorhalfofbranchialregiondevoidoftubercles.................................2011.Urogastricregionwithawidetubercle;mesogastricandprotogastricregionswith6prominenttuberclesarrangedina‘Y’.........................H.pleione(Herbst,1803)–Urogastricregionsmooth;mesogastricandprotogastricregionswithlowtuberclesornomorethan4prominenttubercles...................................................1212.Branchialsubmarginwithatubercleabovechelipedbaseand2-5submarginaltuberclesbehindit................................................H.hilgendorÞDeMan,1887–Branchialsubmarginwithatubercleabovechelipedbasebutnoothertubercles........1313.Merusofpereopods2and3with2or3spinesonextensormargin..........H.barun.sp.–Merusorpereopods2and3withoutspinesonextensormargin.......................1414.Postorbitallobelong,slender...H.ternatensisBuitendijk,1939–Postorbitallobeshort,broad.......................................................1515.Basalantennalarticlewithadeepconcavityonthemarginbehindtheanterolateralspine.Lateralprotogastrictuberclegenerallylow..........................................16–Basalantennalarticlewithashallowconcavityonthemarginbehindtheanterolateralspine.Lateralprotogastrictuberclegenerallyprominent....................................1716.Intestinalregionwithamedialspine........

8 ..............H.subinermisZehntner,1894–
..............H.subinermisZehntner,1894–Intestinalregionsmoothorwithalowtubercle..................H.convexusMiers,188417.Rostralspines1/2to2/3postrostralcarapacelength;gastricregionsstronglyelevatedwith2prominentmesogastrictubercles;branchialregionwithastronglateralbranchialspinedirectedlaterally...............................H.ambonensisGrifn&Tranter,1986–Rostralspines2/3to11/3postrostralcarapacelength;gastricregionsweaklytomoderatelyelevated,mesogastrictubercleslowordistinctbutnotprominent;branchialregionwithaveryshortepibranchialspine,sometimescurvedforwards............................1818.Hepaticmarginandintestinalregioneachwithaspine;anteriorbranchialregionwith1dorsaltubercle..............................................H.unciferCalman,1900 AMANDAM.WINDSOR&SHANET.AHYONG–Hepaticmarginandintestinalregionwithatubercle;anteriorbranchialregionwith2dorsaltubercles........................................................................1919.Preorbitalspineofeavehorizontal;anteriormarginofpostorbitallobeweaklysinuous.Firstpleopodofmale(G1)withamediallobebehindapexH.borradailei(Rathbun,1907)–Preorbitalspineofeaveslightlyupturned;anteriormarginofpostorbitallobenearlystraight.Firstpleopodofmale(G1)withalaterallobebehindtheapex...H.brockiiDeMan,188720.Rostralspinesveryshort;1/6to1/4pcl.............H.planasius(Adams&White,1848)–Rostralspinesatleast1/3,moreusually1/2pclormore..............................2121.Mesogastricregionwith1-3tuberclesorspines.....................................22–Mesogastricregionsmooth........................................................2622.Branchialregionwithanepibranchialspine.........................................23–Branchialregionwithanepibranchialtubercleorsmoothbutnotwithaspine..........2523.Mesogastricregionwithalargetubercleorspineatthesummitandaspineorlargetubercleontheanteriorslope..............................H.spinosusA.Milne-Edwards,1872–Mesogastricregionwithatubercleatthesummit,anteriorslopesmoothorwithaverylowtubercle.........................................................................2424.Rostralspinessubparallel;carapacena

9 rrowposteriorly(cw0.6pcl)...............
rrowposteriorly(cw0.6pcl)...................H.espinosus(Borradaile,1903)–Rostralspinesdivergent;carapacebroadposteriorly(cw0.7-0.75pcl).................H.diacanthus(DeHaan,1839)25.Rostralspinesparallelorweaklydivergent(distancebetweentipsabout11/2timesbasalwidth).Anterolateralangleofbasalantennalarticlenotproduced.......................H.elongatusOrtmann,1893–Rostralsinesstronglydivergentfrombase(distancebetweentipsabout3timesbasalwidth).Anterolateralangleofbasalantennalarticlewithaslenderspine........................H.kyusyuensis(Yokoya,1933)26.Anterolateralangleofbasalantennalarticlesharp,withashorttomoderatespine,sometimesbid............................................................................27–Anterolateralangleofbasalantennalarticlenotproduced,rounded,blunt..............3027.Preorbitalangleofeaveproducedfurtherlaterallythanalmostobsoleteantorbitalangle...H.bispinosusBuitendijk,1939–Preorbitalandantorbitalanglesofeaveaboutequallyproducedlaterally...............2828.Rostralspinesabout2/3pclormore;hepaticregionwithasmalltubercle....H.cracentisGrifn&Tranter,1986–Rostralspines1/2pclorless;hepaticregionsmooth.................................2929.Antorbitalangleofeaverounded.Anterolateralangleofbasalantennalarticlewith1spine.Epibranchialspinelackingorverysmall....................H.inermis(Rathbun,1911)–Antorbitalangleofeaveacute.Anterolateralangleofbasalantennalarticlewith2spines.Epibranchialspinesmallbutdistinct.......................H.minutusBuitendijk,193930.Epibranchialspinerobust,about1/6to1/2cw.Teethondactylofpereopod5verysmall..H.campbelliGrifn&Tranter,1986–Epibranchialspineshortabout1/12cw;teethondactylofpereopod5robust.....H.hendersoni(Laurie,1906) HYASTENUSBARUNOV.ACKNOWLEDGEMENTSWearegratefultoAlainCrosnierfortheloanofspecimensandforhishospitalitytoSTAduringvisitstotheMuséumnationald’Histoirenaturelle.AmandaWindsor’sresearchattheAustralianMuseumwassupportedbyanEndeavourFellowshipfromtheAustralianGovernment(No.3295_2012).,W.T.,1909.OndecapodCrustaceafromChristmasIsland,collectedbyDr.C.W.Andrews,F.R.S.,F.Z.S.ProceedingsoftheZoologicalSocietyofLondon

10 :703-713.,J.G.,1888.BerichtüberdievonHer
:703-713.,J.G.,1888.BerichtüberdievonHerrnDr.J.BrockimIndischenArchipelgesammeltenDecapodenundStomatopoden.ArchivfürNaturgeschichte,(1):289-600,pls.11-228.RIFFIN,D.J.G.&H.A.T,1986.TheDecapodaBrachyuraoftheSibogaExpedition.PartVIII:Majidae.Siboga-Expeditie,:1-335.,L.W.&P.K.L.N,1999.NotesonsomesoutheastAsianspeciesofspidercrabsofthe(Crustacea:Decapoda:Brachyura:Majidae).RafesBulletinof(1):59-72.,P.K.L.,D.GUINOT&P.J.F.DAVIE,2008.SystemaBrachyurorum:PartI.Anannotatedchecklistofextantbrachyurancrabsoftheworld.RafesBulletinofZoology,ATHBUN,M.J.,1916.NewspeciesofcrabsofthefamiliesInachidaeandParthenopidæ.ProceedingsoftheUnitedStatesNationalMuseum,:527-559.,L.,1894.VoyagedeMM.M.BedotetC.Pictetdansl’ArchipelelMalais.Crustacésdel’ArchipelMalais.RevueSuissedeZoologieetAnnalesduMuséed’HistoireNaturelledeGenève,:135-214,pls.VII-IX.Comparativematerial.—HyastenusambonensisGrifn&Tranter,1986:AMP34609,1spentfemaleparatype(cl20.0mm,pcl14.6mm,cw9.3mm),AmbonBay,Ambon,90m,dredge,stones,sand,DanishKeiIslandsExpedition,3Mar1922.—Hyastenusborradailei:AMP13893,2males(cl9.2mm,pcl6.1mm,cw4.0mm;cl16.2mm,pcl8.8mm,cw5.5mm),NinetyMileBeachbetweenCapeJaubertandWallal,WesternAustralia,9m,dredge,coll.A.A.Livingstone,Sep1929;AMP24104,1male(cl19.8mm,pcl9.3mm,cw6.1mm),1female(cl19.4mm,pcl9.0mm,cw5.9mm),BroadhurstReef,Townsville,Queensland,onhydroid(?sp.),10.5m,coll.I.Loch,11Oct1975.—HyastenusbrockiiDeMan,1887:AMP34617,1female(cl11.6mm,pcl7.3mm,cw4.9mm),AmbonReef,Ambon,SibogaExpedition;AMP34618,1female(cl11.6mm,pcl6.4mm,cw4.0mm),Banda,offWaling,15m,sand,DanishKeiIslandsExpedition,15May1922;AMP34616,1male(cl10.9mm,pcl6.2mm,cw4.1mm),bankbetweenBahuluwangandTambulungenislands,Salayer,southSulawesi,8-10m,dredge,deadcoral,,lithithamnion,Sibogasta.66,7-8May1899.—HyastenussubinermisZehntner,1894:AMP34639,1male(cl12.0mm,pcl8.1mm,cw5.6mm),1ovigerousfemale(cl16.1mm,pcl11.4mm,cw8.1mm),Ambonreef,Ambon,SibogaExpedition.—HyastenusunciferCalman,1909:AMP34640,1male(cl11.7mm,pcl5.7mm,cw3.6mm),offNeiraBanda,13m,sand,DanishKeiIslandsExpedition,coll.Dykker,12Jun1922.Firstreceived20December2012.Finalversionaccepted9March2