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Anal Bot Cavanille 3 (1975TH SPANIS GENTIANby Askell LOVE and Doris LO Anal Bot Cavanille 3 (1975TH SPANIS GENTIANby Askell LOVE and Doris LO

Anal Bot Cavanille 3 (1975TH SPANIS GENTIANby Askell LOVE and Doris LO - PDF document

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Anal Bot Cavanille 3 (1975TH SPANIS GENTIANby Askell LOVE and Doris LO - PPT Presentation

U2 ANALE DE INSTITUT BOTANIC A J CAVANILLES TOM Ino such epithets were available A preliminary review of the situation an o th basi fo th reclassificatio ha recentl bee give bLov LovSinc 196 w hav ID: 283886

AN'ALE INSTITUT BOTANIC

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Anal Bot Cavanille 3 (1975TH SPANIS GENTIANby Askell LOVE and Doris LOVE Dedicated to Prof. S. Rivas Goda o anniversarAlthoug th classificatio int a singl genu o th mor tha60 U2 AN'ALE DE INSTITUT BOTANIC A J CAVANILLES TOM Ino such epithets were available. A preliminary review of the situation an o th basi fo th reclassificatio ha recentl bee give bLov & LovSinc 196 w hav ha severa occasion t stud th gentian oth Iberia Peninsula ai o a generou NAT Scienc GranNo (cf Lov & Kjellqvis 1972 1973 1974a b an late witothe an indirec suppor o durin shor visit a tourists Becausth cytologica result ar connecte wit th taxonomica conclusiono th generi designatio o th group i question thes ar reportebelo unde th vali nomenclatura combination tha 'w believ ought replac olde name base o th traditionall collectiv genericoncept1. Gentiana L., s. str. When the collective genus Gentiana is split up into more natural units Linnaea genu Gentian mus b restricte t th sectioCoelanth o Kusnetzo sinc thi sectio i a somewha heterogeneou assemblag o th typ specie G lute L (cfHitchcoc & Greene i Brique 1935 whic automaticall constituteth sectio o subgenu Gentian i th genu i s subdivided anth othe fou o fiv specie whic diffe fro i i flower morphologan perhap eve i polle typ (Nilsso wa conclude bHolu (1970 tha th latte ough t b place i a distinc subgenuCoilanth (Borckh. Holub Hybrid betwee al th specie o Coilanth s restricte see t b commo whereve thes tax mee anfor hybri swarm throug backcrossin t eithe parent s theireproductiv isolatio fro eac othe i questionable Furthermorehybrid occu betwee G lute an th othe species a ha beeknow fo a considerabl tim (cf Fock thes see t bpartiall fertile Therefore w doub th wisdo o separatin thestax i differen section o subgenera -wherea ther seem t b a reaso fo wonderin i the migh no rathe b reduce t subspecifi o varieta statu withi tw genu i characterizeb it larg 5-7-merou yello o purpl a rotat an deeplparte corolla sometime wit plait whic the ar smal an asymmetrical seed ar broadl winged an th plant ar perenniawit broa an petiolat leaves TH SPANIS GENTIAN22Th genu Gentian i th stric sens i cytologicall characterizeb th basi numbe x = 5 al th tax ar tetraploi wit 2 n — 4chromosomes thei karyotyp i similar perhap wit a mino•deviatio i th frequenc o exactl metacentri chromosome iG lutea Th followin tw specie ar represente i SpainGentiana lutea L. 2 n = 40, determined on material of subsp. lutea fro nea Gofi i th Sierr d Andi wes o Pamplona voucheG-Olfll Thi confirm previou report b Favarge (1949Gentian Burser Lapeyr 2 = o plant belongin tth subsp Burser fro nea Setcasa i th Sierr d l Fembr Morti th Pyrenees vouche G-0811 confirmin previou report b Kupfe& Favarge (1967 an Favarge & Kupfe (1968 als fro thPyrenees2. Pneumonanthe Gleditsch Another group, which is well defined morphologically, palynologically an cytologically i forme b th sectio Pneumonanthe provide thath specie G asclepiade L an it clos relativ G schistocaly CKoc ar exclude a th separat genu Dasystephan Adans (cf DLove Nilsson 1967 Lov & Love genu stricte i characterize b it lac o a basa lea rosette smal anscale-lik lowe leaves a limbles coroll wit erec lobe an weldevelope plaits ofte connat anthers an usuall bu no alwaywinge seeds I share a simila polle morphology see coa structure th basi numbe x = 1 wit th followin genus o whici migh therefor b regarde a a subgenu (Lov & LovHowever sinc w ar no awar tha th specie o thes tw groupeve hybridiz i natur o experimentally despit ou effort includinvariou tax o both w find i mor logica t retai the a distincgenera followin th principl tha i i wise an evolutionarilmor correc t separat tax tha ma b simila tha t unit thoswhic ar possibl distinct Th typ specie G pneumonanth L wagive a generi statu a Pneumonanth b Gleditsc Dand1967) onl Spanis specie o tha genu s restricte isPneumonanth vulgari F W Schmidt 2 n = 26 determine o a collectio fro nea Gofi i th Sierr d Andi wes o Pamplona 22 ANALE DE IKSTIIUT BOTANIC A J CAVANILLES TOM Ivoucher G-0397. This confirms earlier reports by several authors (cf & Lov3. Tretorhiza Adans. This group, or the section Cruciata Gaudin, which Kusnetzov (1 rename sectio Aptera i simila t th las genu i it pollemorphology see coa structur an basi chromosom number bui differ fro i morphologicall b it crow bein envelope ifibrillos sheath o ol leaves an i tha th lowe leave for a rosette coroll i limbles o obscurel limbed th plait ar largan usuall symmetrica an biparted th anther ar fre an th seedar wingless Th onl representativ o thi genu i Spai i thtyp • Tretorhiza cruciata (L.) Opiz. 2» = 52, determined on material fro nea Sardena i th Sierra d Sa Gra wes o Figuerasvouche G-OJ^IS Th sam chromosom numbe ha bee reportepreviousl b severa author (cf Lov & Lov4. Ciminalis Adans. The section Megalanthe Gaudin, or the section Thylacites Griseb. of mos manuals wa tentativel unite wit th sectio ChondrophyllBge th nam Ericoil Ren b Lov & Lov (1961ain th validatio o thi pre-Linnaea nam b BorckhauseThi wa premature no onl becaus th nam Ericoil i th circumscriptio give t i b Borckhause an eve b Gra (1821unde th correcte spellin Erical S F Gray mus b rejecte fovariou reason (cf Holu als becaus th smal grouo perennia specie o Chondrophyll wit th basi chromosomnumbe x = 3 i unrelate t th sectio Megalanth an t th larggrou o annua o biennia specie wit x = 9 traditionall includei th sectio Chondrophylla Th perennial o th sectio Megalanth an th annual-biennia specie o Chondrophyll are howevermorphologicall close an als know t b abl t hybridiz i naturan unde experimenta conditions Therefore w believ tha thear correctl accomodate i a singl genus a conclude b Lov & TH SPANIS GENTIAN225Lov (1961a correc namGentian acauli (cf genurepresente thCiminalis acaulis (L.) Moench. 2 n = 36, determined on plants from nea Gon th wes Vouche G-0^08Thi confirm earlie report (1949LovkCiminalis alpina (Vill.) Holub. 2 n = 36, determined on plants from nea Setcasa th l Mort thVouche G-0311 Thi confirm earlie report (1949Favarge & KupfeCiminalis occidentalis (Jakowatz) Holub. 2 n = 36, determined on materia fro belo th Pic Bassagod wes o Figuera i th easterPyrenees G-0)$k Thi confirm a previou repor fro thPyrenee b Favarge & Kupfe (1968 unde th synony G angustifoli5 Holubi Lov & LoveAs mentioned in connection with the genus Ciminalis, the perennial specie o th sectio Chondrophyll ar morphologically cytologicallan eve palynologicall distinc fro th annua specie traditionallclassifie i thi section wherea th latte ar congeneri wit th taxusuall groupe i th sectio Megalanth fo whic th correc generinam i Ciminali Adans Fo thi smal perennia group Lov & Lov (1975 use provisionall th generi nam Erical S F Grayhowever tha nam mus b regarde a bein illegitimat o synonymou wit Ciminali an must therefore b rejected Sinc tha leaveth perennia specie o sectio Chondrophyll withou a legitimat nama th generi level althoug the certainl constitut a natura anwell-define genus a ne nam mus b furnishe t accomodat thesabou te specie o Eurasia Nort an Sout Americ wit th basichromosom numbe x = b th specie Gentian pyrenaic L Fo thi grou w propos th name •22 ANALE DE INSTITUT BOTANIC A J CAVANILLES TOM IHolttbia Love & Love, now gen. Planta perennis, rhizomata repente ramoso caudiculis plurimis epi-geis firmis erecti 3-1 c alt. foli acuta subobtus vemucronata coriace marginata calyc campanulat nodentato tubuloso-campanulata tubuloso-clavat ve infundibularo-campanulata ovari oblongo stigmatibu ovatis Chromosomatx = 13A perennia plan wit creeping branchin an slende rootstocwit numerou epigeou offshoots stem firm an c longleave acute subobtus o mucronate coriaceous caly campanulateunsplit coroll tubular-campanulate tubular-clavate o campanulate-infundibular ovar oblong style ovate Basi chromosomnumbe x = 13Typ specie o th genu i Holubi pyrenaic (L. Lov & Love•comb base o Gentian pyrenaic L. MantissTh chromosom numbe 2 n =2 ha bee determine fo Holubipyrenaic fro th Sierr d l Cafi nort o Riba d Fresse i th-easter Pyrenees Vouche G-O^Jfi I confirm earlie report bSokolovskay & Strelkov & Favarge (1967 anFavarge & KupfeAnother Spanish species of the genus is: Holubia boryi (Boiss.) Love & Love, comb, nov., based on Gentiana bory Boissier Biblioth Univ Geneve ser 2,1 It•chromosom number whic ha no bee previousl reported i2 =2 a counte o materia fro nea Pto d l Bonaigu i th•centra Pyrenees Vouche G-0522(!. Calathiana Delarbr The section Cyclostigma Griseb. or section Calathianae Froelich include abou te perennia o rarel annua specie wit leave thaar sometime i a rosette an wit shor stem usuall wit a singltermina flower. Th caly i herbaceous unspli bu wit five teethTh corolla-tub i almos cylindrica an plicate bu th coroll itseli blu an hypocraterifor wit paten lobe an a smal appendagi th anther ar th stigma-lobe ar contiguouan fuse int a circula an slightl concav dis wit dentate-fimbriat TH SPANIS GENTIAN22margins capsul i sessil an th seed ar no winged Thbasi chromosom numbe seem t b x = 7 o whic diploid tetraploi an hexaploi multiple ar known th basi number x = 8 an1 hav als bee reported Ther seem t b n palynologica oothe indicatio o heterogeneit withi th group whic i als characterize b a homogenou see coa structure s ther i perhap a reaso t wonde i th chromosom numbe report deviatin froth basi numbe x — 7 migh hav bee cause b th occurrenc oB-chromosomesLov & Lov (1961a b identifie thes plant o high-alpin an:arcti region i th borea zon preliminaril wit th genu HippioF W Schmidt Recen studie have however reveale tha thi nam-mus b regarde a illegitimat an tha th oldes correc namavailabl fo thi taxo a th generi leve i Calathian Delarbr•(Holu th typ spece C nivali (L. Delarbre Th genui represente i Spai b th specieCalathian brachyphyll (Vill. Holub 2 = counte oplant fro nort o Estaho i th Sierr d Campirm i th centraPyrenees G-04%9 Thi verifie earlie report b Favarge& Huyn (i Lov & Solbri 1964 an FavargeCalathian delphinensi (Beauverd Holub 2 n = oplant fro nea th Pic Bassagod wes o Figuera i th easterPyrenees G-OJ/29 Thi i a ne numbe fo thi taxo oth S Alp an th easter Pyrenees whic Fourniea a subspecie onl o th otherwis easter Alpiqu specie Gentianpumil Jacq ( Ciminali pumil (Jacq. easter Alpiqutaxo 'wa reported b Favarge hav th chromosomnumbe 2 « = 20 whic ma o ma no hav bee determine o a hybri individua (2 = 21?)Calathian nivali (L. Delarbre 2 n = o materiafro nea Setcasa i th Sierr d l Fembr Mort i th PyreneesVouche G-0309 Thi numbe ha previousl bee reporte fro•elsewher i th Pyrenee b Favarge & Kupfe (1968 an b severaauthor fro othe region (cf Lov & LovCalathian vern (L. Holub 2 n = o materiaginatin fro th slope o Pic Bassagod nort o Sardena i theaster Pyrenees Vouche (7-0.J8S Thi confirm earlie count bsevera author (cf Lov & Lov 22 ANALE DE IKSTITUT BOTjta'IC A J CAYANILLES TOM I7. Comastoma Toyokuni. The section Comastoma Wettst. is often placed in the collective genu Gentianella o whic i ha sometime bee assume t b themos primitiv species althoug i i apparentl no closel relate toft bu instea ma b neare t Lomatogoniu A Brau wit whic ishare certai morphologica features th basi chromosom numbex = 5 an somewha simila polle morphology Lov & Lovconclude fro thi resemblanc observe alread b Wettstei (1806tha th sectio Comastom ough t b transferre t Lomatogoniumwherea Toyokun maintaine tha Comastom i actuallmor correctl place i a genu o it own A such thi taxoninclude les tha twent species whic ar know t occu mainl irth easter Asiati mountains althoug som tax ar als representei Europ an Nort Americ an i th Arctic Th specie ar annual o biennial usuall wit long-pedicelle flowers, a shor calyxtube o campanulat coroll tha i 4 o 5-merou anha on o tw fimbriate scale a th bas o eac lobe ther i nostyl an onl ver shor stigma an th seed ar Spainthi genu i represente b th speciesComastoma tencllum (Rottb.) Toyokuni. 2 n = 10, as determined on materia fro nea Pto d l Bonaigu i th centra Pyrenees Vouche G-0519 Thi coun confirm earlie report b severa authorsfro elsewher (cf Lov & Lov8. Gentia'nopsis Ma. Another section of Gentiana that some authors still include as * par o th genu Gentianella i th sectio Crossopetalu Froel Ideviate considerabl fro al th othe group i genera an floral morpholog (Lindsa 1040 bu als i polle morphology see coastructur an i cytologica characteristics a pointe ou b D Love(1053 an Nilsso i i bes characterize bit Avit a unspli caly an a tubular-infundibulacoroll withou a fringe rin i th throat bu th coroll lobe arfrequentl fringe o ciliat a th margin Th styl i distinc an T1I SIWN'IS GENTIAN22ova o tubular an th capsul i stipitate Th basi chromosomnumbe i x = 11 Th grou include abou 7 annua o bienniaspecie o mainl Asiati an Nort America distribution thoug i ials represente b a fe specie i alpin an subarctic-arcti EuropeA th generi level th taxo ha bee give th invali nam An.thopogo Necke (cf Stafle & ali 1972 an als th nam Crossopetalu Roth whic ha a earlie homony i th Celastraceaewherea it vali nam i Gentianopsis a describe b MSpai i i represente b a singl speciesGentianopsis ciliata (L.) Ma. 2 n = 44, as counted on material originatin fro nort o Estaho i th Sierr d Campirm i th•centra Pyrenees Vouche G-0-'f%7 Thi confirm previou report bFavarge (1949 (1957 an Skalixsk (i Skalinsk & ali9. Gentianella Moench. The genus Gentianella in its strict sense as typified by Gentianella •campestri (L. Borne include th Gentian section Endotrich Froelan Arctophil Griseb o borea mountains I comprise abou 10 annua o biennia specie wit 4-5-merou o cylindricacoroll whic i th throa an wit erec o paten lobesn sessil an stipitat capsule I ha th basi chromosom-numbe x = 9 an a polyploi serie fro th diploi t th hexaploidI Spain thi genu s restricte i represente b tw speciesGentianella campestris (L.) Borner. 2 n = 36, determined on mate­ria collecte o th nort slope o Pic Bassagod i th easterPyrenees G-0^05 Tha verifie earlie report b Favarge(1949 fo th typica subspecie an b D Lov (1953 an Lov & Lov (1956 fo th subsp islandicaGentianella hypericifolia (Murb.) Pritchard. 2 n = 36, counted on plant fro th sout slope o Mont Ad nort o Cilbet i th westerPyrenees G-0381 Thi confirm earlie report b Favarge & Huyn (i Lov & Solbri & Favarge (1967an Favarge & Kupfe 23 ANALE DE INST1TUT BOTANIC A J CAVANILLES TOM IR E S U M E N Al reclasiricar las gencianas espanolas en un sistema mas natural que l agrupacio tradicional com seccione de gener colectiv Gentiana pertenece a lo genero restringido Gentian L. s str.Pneumonanth Gleditsch Thetorhis Adans. Ciminali Adans. Calathian Delarbre Comastom Toyokuni Gentianopsi Ma GentianellMoenc y Holubi Lov & Love describiendos est ultim e e present trabaj par inclui la especie perenne d l tradicionalmentdelimitad seccio Chondrophylla Tambie s relaciona la especiecuy presenci e Espan e conocid d form natural co indicaciod lo ntimero cromosomico determinado sobr materia espaiiolAbstracWhen reclassified into a more natural system than the traditional" groupin a section onl o th the ver collectiv genu Gentiana thSpanis gentian belon t th restricte gener Gentian L. s str.Pneumonanth Gleditsch Tretorhiz Adans. Ciminali Adans. Calathian Delarbre Comastom Toyokuni Gentianopsi Ma GentianellaMoench an Holubi Lov & Love th las on bein describe i thipape t accomodat th perennia specie o th traditionall delimitedsectio Chondrophylla Th specie know t occu naturall i Spaiar th following wit chromosom number determine o Spanishmateria give i parentheses Gentian lute L subsp lute (2 =Gentian Burser Lapeyr subsp Burser ( n =vulgari F W Schmid ( n = cruciat (L. Opi( = acauli (L. Moenc ( n = alpin(Vill. Holu ( n = occidentali (Jakowatz Holub( w = pyrenaic (L. Lov & Lov ( m =Bory (Boiss. Lov & Lov (2 = brachyphyll (Vill.)Holu ( n = delphinensi (Beauverd Holu ( n = = nivali (L. Delarbr ( n = vern(L. Holu = tenellu (Rottb. Toyokun ( re = = ciliat (L. M ( n = campestris(L. Borne ( n = h\pericifoli (Murb. Pritchart( = 36) TH SPANIS GENTIAN23BibliographBorckhausen, D. B. M. — 1796—Ueber Linnes Gattung Gentiana — Roemer: Arch-Bot.Briquet — Internationa rule o botanica nomenclatur — JenaDandy J E —196 — Inde o generi name o vascula plant 1753-177 — RegnunVegetabile 51 1-130F'avarger C —194 — Contributio a e biologiqu de Gentianacees Bot Ges. 59 62-86— — 195 — Contributio a e biologiqu de Gentianacees I— Bot Ges. 62 244-2571959—Note d caryologi Alpine Il — Neuchat Sci Nat. 82:255-285-196 — Note d caryologi Alpine I — Neuchat Sci Nat. 885-60— — & Kiipfer P —196 — Contributio a l cytotaxonomi d l flor alpin desPyrenees Collectane Botanica 7 325-357Focke O — 188 — Di Pflanzen-Mischling — BerlinFournier P —1938—Le quatre Flore d l France Cors compris — Poinsonles-Grancey pp 833-896Gleditsch J G — 176 — System naturu a staminu BerlinGray S F — 182 — A natura arrangemen o Britis plant — LondonHolub 197 — Ne name i Phanerogamae 1 — Foli Geobot Phytotax. 5435-441197 — Ne name i Phanerogamae 2 — Foli Geobot Phytotax. 8 155-179Ktipfer P & Favarger C —196 — Premiere prospectu caryologique dan 1flore orophil de Pyrenee e d l Sierr Nevad — C r Acad Sci Paris 2642463-2465Kusnetzov N — 189 — Gentian Tournef — Pflanzenfam. 4 (2)80-86• 1896-190 — Subgenu Eugentian Kusnetz generi Gentian ActHort Petrop. 15 1-507Lindsay A A —194 — Flora anatom i th Gentianacea — Amer Journ Bot.,27Love —1971—IOP chromosom numbe report XXXI —Taxo 20 808-821& Kjellqvist E.—1972—Cytotaxonom o Spanis plants I Introduction.Pteridophyt an Gymnosperma — 23-35— — & Kjellqvist E — 197 — Cytotaxonom o Spanis plants II Monocotyledon — 147-182& Kjellqvist E — 1974 — Cytotaxonom o Spanis plants III Dicotyledon — 3-32— — & Kjellqvist E — 1974 — Cytotaxonom o Spanis plants IV DicotyledonsCaesalpiniaceae-Asteraceae—Lagascalia 4 153-211& Love D — 195 — Cytotaxonomica conspectu o th Icelandi ActHort Gotob. 20 65-291 123 ANALE DE INST1TUT BOTANIC A J CAYANIIXES TOM XXXII VOL I• & Love, D. — 1961a — Some nomenclatural changes in the European flora. I. Specie an supraspecifi categorie — 114& Love D — 1961 — Chromosom number o Centra an Northwes Europeaplan specie — Oper Botanica 5& Love D —197 — Cytotaxonomica atla o th Slovenia Lehre& Love D — 197 — Th natura gener o Gentianacea — Recen Adv iBotany P N Mehr Jub. vol. (i& Solbrig O T — 196 — IOP chromosom numbe reports I — Taxon13Love —195 — Cytotaxonomica remark o th Gentianacea —225-235Ma — 195 — ne genu o Chines Gentianacea — Act Phytotax Sinic —Nilsson S —196 — Polle morphologica studie i th Gentianaceae-Gentianina — Gran Palynologica 7 46-145Quezel —195 — Peuplemen vegeta de haute montagne d Nor— Encyclop Biogeogr Ecol. 10 1-445Rork C L.—1949— Cytologica studie i Gentianacea — Bot. 36687-701Skaliriska M. Czapik R. Piotrowicz M & alii.—195 — Furthe studie i chromosom number o Polis angiosperm Act Soc Bot Polon.28Sokolovskaya A P & Strelkova O raspredeleni polyploidov III Issledovani flory alpiysko oblast tsentralnog kavkazkog khrebt— Uchen Zap Ped F A Inst Im Gertsena C6 195-216Stafleu F A & — Internationa cod o botanica nomenclatur — UtrechtToyokuni H.—196 — Separatio d Comastotna genr nouveau— Tokyo 74 198196 — Furthe remark o th genu Comastotn — Act Phytotax Geobot.20Wettstein R — Di Gattungszugehorigkei un systematisch Stellun deGentian tenell Rottb un G nan Wulf — Zeitschr. 46 172-176Institute of Alpine and Arctic Research Universit o ColoradBoulder Colorad 80302 U S A