Posterior cavity Iris Ciliary body Choroid Cornea Sclera Neural part Pigmented part Anterior cavity Fibrous layer Vascular layer uvea Inner layer retina Horizontal section of right eye ID: 742132
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Slide1
Figure 17-5b The Sectional Anatomy of the Eye.
Posterior
cavity
Iris
Ciliary
body
Choroid
Cornea
Sclera
Neural part
Pigmented part
Anterior
cavity
Fibrous
layer
Vascular layer
(uvea)
Inner layer
(retina)
Horizontal section of right eye
bSlide2
Figure 17-5a The Sectional Anatomy of the Eye.
Fovea
Retina
Choroid
Sclera
Sagittal section of left eye
Corneal
limbus
Iris
Pupil
Cornea
Ora
serrata
Ocular conjunctiva
Eyelash
Lens
Fornix
Palpebral conjunctiva
Optic
nerve
aSlide3
Figure 17-6 The Pupillary Muscles.
Pupillary constrictor
(sphincter)
Pupil
Pupillary dilator
(radial)
The
pupillary dilator
muscles
extend radially
away from the edge of the pupil.
Contraction of these muscles
enlarges the pupil.
The pupillary constrictormuscles form a series ofconcentric circles around thepupil. When these sphinctermuscles contract, the diameterof the pupil decreases.
Increased light intensityIncreased parasympathetic stimulation
Decreased light intensity
Increased sympathetic stimulationSlide4
Figure 17-7a The Organization of the Retina (Part 1 of 2).
Amacrine
cell
Horizontal cell
Cone
Rod
Pigmented
part of retina
Rods and
cones
Bipolar cells
Ganglion cells
LIGHT
The cellular organization of the retina. The photoreceptors are closest to the choroid, rather than near the posterior cavity (vitreous chamber).
aSlide5
Figure 17-7b The Organization of the Retina.
The optic disc in diagrammatic sagittal section.
Optic disc
Neural part
of retina
Pigmented
part of retina
Central retinal vein
Central retinal artery
Optic nerve
Sclera
Choroid
bSlide6
17-3 The EyeHorizontal and
Amacrine Cells Facilitate or inhibit communication between photoreceptors and ganglion cellsAlter sensitivity of retinaOptic DiscCircular region just medial to foveaOrigin of optic nerveBlind spotSlide7
17-3 The EyeThe
LensLens fibersCells in interior of lensNo nuclei or organellesFilled with crystallins, which provide clarity and focusing power to lensCataractCondition in which lens has lost its transparencySlide8
17-3 The EyeLight
RefractionBending of light by cornea and lens Focal pointSpecific point of intersection on retinaFocal distanceDistance between center of lens and focal pointSlide9
Figure 17-10 Factors Affecting Focal Distance.
Focal distance
Close
source
Focal distance
Focal
point
Lens
Light
from
distant
source
(object)
Focal distance
The closer the light source,
the longer the focal distance
The rounder the lens,
the shorter the focal distance
b
c
aSlide10
17-3 The EyeLight Refraction of Lens
AccommodationShape of lens changes to focus image on retina AstigmatismCondition where light passing through cornea and lens is not refracted properly Visual image is distortedSlide11
Figure 17-11 Accommodation.
a
b
Lens rounded
For Distant Vision:
Ciliary
Muscle Relaxed, Lens Flattened
For Close Vision:
Ciliary
Muscle Contracted, Lens Rounded
Ciliary
muscle
contracted
Focal point
on fovea
Lens flattened
Ciliary
muscle
relaxedSlide12
17-3 The EyeLight Refraction of Lens
Image reversalVisual acuityClarity of vision“Normal” rating is 20/20Slide13
Figure 17-12a Image Formation.
a
Light from a point at the top
of an object is focused on
the lower retinal surface.Slide14
Figure 17-12b Image Formation.
b
Light from a point at the
bottom of an object is focused on the upper
retinal surface.Slide15
Figure 17-12c Image Formation.
c
Light rays projected from a vertical
object show why the image arrives
upside down. (Note that the image
is also reversed.)
Optic
nerveSlide16
Figure 17-12d Image Formation.
d
Light rays projected from a horizontal
object show why the image arrives with
a left and right reversal. The image also
arrives upside down. (As noted in the
text, these representations are not
drawn to scale.)
Optic
nerveSlide17
Figure 17-13 Refractive Problems (Part 1 of 5).
The eye has a fixed
focal distance and
focuses by varying theshape of the lens.
A camera lens has a
fixed size and shapeand focuses by varying
the distance to the film.Slide18
Figure 17-13 Refractive Problems (Part 2 of 5).
Emmetropia
(normal vision)Slide19
Figure 17-13 Refractive Problems (Part 3 of 5).
Myopia
(nearsightedness)
If the eyeball is too deep or the resting
curvature of the lens is too great, the
image of a distant object is projected
in front of the retina. Myopic people
see distant objects as blurry and out
of focus. Vision at close range will be
normal because the lens is able to
round as needed to focus the image
on the retina.
Myopic corrected witha diverging,concavelens
DiverginglensSlide20
Figure 17-13 Refractive Problems (Part 4 of 5).
Hyperopia
If the eyeball is too shallow or the lens
is too flat, hyperopia results. The
ciliary muscle must contract to focus
even a distant object on the retina.And at close range the lens cannot
provide enough refraction to focus animage on the retina. Older peoplebecome farsighted as their lenses lose
elasticity, a form of hyperopia calledpresbyopia (
presbys, old man).
(farsightedness)
Hyperopia
corrected with
a converging,convexlens
ConverginglensSlide21
Figure 17-13 Refractive Problems (Part 5 of 5).
Surgical Correction
Variable success at correcting myopia and
hyperopia has been achieved by surgery that
reshapes the cornea. In photorefractive
keratectomy (PRK) a computer-guidedlaser shapes the cornea to exact specifications.
The entire procedure can be done in less thana minute. A variation on PRK is called
LASIK(Laser-Assisted in-Situ Keratomileusis
). In thisprocedure the interior layers of the cornea arereshaped and then recovered by the flap of original
outer corneal epithelium. Roughly 70 percent of LASIKpatients achieve normal vision, and LASIK has become the most
common form of refractive surgery. Even after surgery, many patients still need reading glasses,and both immediate and long-term visual problems can occur.Slide22
17-4 Visual PhysiologyVisual Physiology
Rods Respond to almost any photon, regardless of energy contentCones Have characteristic ranges of sensitivitySlide23
17-4 Visual PhysiologyAnatomy of Rods and Cones
Outer segment with membranous discsInner segmentNarrow stalk connects outer segment to inner segmentSlide24
17-4 Visual PhysiologyAnatomy of Rods and Cones
Visual pigmentsIs where light absorption occursDerivatives of rhodopsin (opsin plus retinal)Retinal synthesized from vitamin ASlide25
Figure 17-14a Structure of Rods, Cones, and Rhodopsin Molecule.
In a cone, the discs are
infoldings
of
the plasma membrane, and the outer
segment tapers to a blunt point.
In a rod, each disc is an independent
entity, and the outer segment forms
an elongated cylinder.
Discs
Connecting
stalks
Mitochondria
Golgi
apparatus
Nuclei
Pigment Epithelium
The pigment epithelium
absorbs photons that are
not absorbed by visual
pigments. It also
phagocytizes old discsshed from the tip of theouter segment.
Melanin granules
Outer Segment
The outer segment of a
photoreceptor contains
flattened membranous
plates, or
discs
, that
contain the visual pigments.
Cone
Rods
Inner Segment
The inner segment contains
the photoreceptor’s major
organelles and is responsible
for all cell functions other
than photoreception. It also
releases neurotransmitters.
Each photoreceptor
synapses with a bipolar cell.
Bipolar cell
LIGHT
Structure of rods and cones
aSlide26Slide27
17-4 Visual PhysiologyColor Vision
Integration of information from red, green, and blue conesColor blindnessInability to detect certain colorsSlide28
Figure 17-21 The Anatomy of the Ear.
Elastic cartilages
External Ear
Auricle
Middle Ear
Auditory
ossicles
Oval
window
Internal Ear
Semicircular canals
Petrous part of
temporal bone
Facial nerve (VII)
Vestibulocochlear
nerve (VIII)
Bony labyrinth
of internal ear
Cochlea
Auditory tube
To
nasopharynx
Vestibule
Round
window
Tympanic
cavity
External acoustic
meatus
Tympanic
membraneSlide29
Figure 17-22a The Middle Ear.
Temporal bone
(petrous part)
Stabilizing
ligaments
Branch of facial
nerve VII (cut)
External
acoustic meatus
Malleus
Incus
Stapes
Auditory
Ossicles
Muscles of
the Middle Ear
Oval window
Tensor tympani
muscle
Stapedius
muscle
Round window
Auditory tube
Tympanic cavity
(middle ear)
Tympanic
membrane
The structures of the middle ear
aSlide30
17-5 The EarVibration of Tympanic Membrane
Converts arriving sound waves into mechanical movementsAuditory ossicles conduct vibrations to inner earTensor tympani muscleStiffens tympanic membrane Stapedius muscleReduces movement of stapes at oval windowSlide31
17-5 The EarThe
Internal Ear Contains fluid called endolymph Bony labyrinth surrounds and protects membranous labyrinthSubdivided into:VestibuleSemicircular canals
CochleaSlide32
Figure 17-23b The Internal Ear.
b
Semicircular
canal
Semicircular ducts
Anterior
Lateral
Posterior
Vestibule
Utricle
Saccule
Vestibular duct
Cochlear duct
The bony and membranous labyrinths. Areas of
the membranous labyrinth containing sensory
receptors (cristae, maculae, and spiral organ) are
shown in purple.
Cristae within
ampullae
Maculae
Endolymphatic
sac
KEY
Membranous
labyrinth
Bony labyrinth
Cochlea
Spiral
organ
Tympanic
ductSlide33
17-5 The EarThe Internal Ear
VestibuleEncloses saccule and utricle Receptors provide sensations of gravity and linear accelerationSemicircular canalsContain semicircular ducts Receptors stimulated by rotation of headSlide34
17-5 The EarThe Internal Ear
CochleaContains cochlear duct (elongated portion of membranous labyrinth) Receptors provide sense of hearingSlide35
17-5 The EarThe Internal Ear
Round window Thin, membranous partitionSeparates perilymph from air spaces of middle earOval windowFormed of collagen fibersConnected to base of stapesSlide36
17-5 The EarStimuli and Location
Sense of gravity and accelerationFrom hair cells in vestibuleSense of rotationFrom semicircular canalsSense of soundFrom cochleaSlide37
17-5 The EarEquilibrium
Sensations provided by receptors of vestibular complexHair cellsBasic receptors of inner earProvide information about direction and strength of mechanical stimuliSlide38Slide39
17-5 The EarThe
Semicircular Ducts Are continuous with utricleEach duct contains:Ampulla with gelatinous cupulaAssociated sensory receptorsStereocilia – resemble long microvilliAre on surface of hair cellKinocilium
– single large ciliumSlide40
Figure 17-24a The Semicircular Ducts.
a
Semicircular ducts
Anterior
Posterior
Lateral
Ampulla
Utricle
Vestibular branch (N VIII)
Cochlea
Endolymphatic
sac
Endolymphatic
duct
Maculae
Saccule
An anterior view of the right
semicircular ducts, the utricle,
and the
saccule
, showing the
locations of sensory receptors.Slide41
Figure 17-24b The Semicircular Ducts.
Ampulla
filled with
endolymph
Hair cells
Crista
ampullaris
Cupula
Supporting cells
Sensory nerve
A cross section through the ampulla of a semicircular duct.
bSlide42
Figure 17-24c The Semicircular Ducts.
Semicircular duct
Endolymph
movement along the length of the duct
moves the
cupula
and stimulates the hair cells.
c
Cupula
Direction of
duct rotation
Direction of relative
endolymph
movement
Direction of
duct rotation
At restSlide43
Figure 17-24d The Semicircular Ducts.
Displacement in
this direction
stimulates hair cell
Displacement in
this direction
inhibits hair cell
Kinocilium
Hair cell
Gelatinous
material
Stereocilia
Sensory nerve
ending
Supporting cell
A representative hair cell (receptor) from the
vestibular complex. Bending the
sterocilia
towardthe kinocilium depolarizes the cell and stimulatesthe sensory neuron. Displacement in the oppositedirection inhibits the sensory neuron.
dSlide44
17-5 The EarThe Utricle and
Saccule Provide equilibrium sensationsAre connected with the endolymphatic duct, which ends in endolymphatic sac Slide45
17-5 The EarThe Utricle and
Saccule MaculaeOval structures where hair cells cluster StatoconiaDensely packed calcium carbonate crystals on surface of gelatinous massOtolith (ear stone) gelatinous matrix and statoconiaSlide46
Figure 17-25ab The
Saccule
and Utricle.
Endolymphatic
sac
Endolymphatic
duct
Utricle
Saccule
The location of
the maculae
a
Otoliths
Gelatinous layer
forming
otolithic
membrane
Hair cells
Nerve fibers
The structure of an individual macula
bSlide47
Figure 17-25c The
Saccule
and Utricle.Head in normal, upright position
Gravity
1
2
Receptor
output
increases
Head tilted posteriorly
Otolith
moves
“downhill,”
distorting hair
cell processes
Gravity
A diagrammatic view of
utricular macular functionwhen the head is held normally and then tiltedback
1
2
cSlide48
17-5 The EarHearing
Cochlear duct receptors Provide sense of hearingSlide49
Figure 17-27a The Cochlea.
Scala
vestibuli
Cochlear duct
Scala
tympani
Cochlear
branch
Vestibular
branch
Vestibulocochlear
nerve (VIII)
Round window
Stapes at
oval window
From oval window
to tip of spiral
From tip of spiral
to round window
KEY
Semicircular
canals
The structure of the cochlea
aSlide50
Figure 17-27b The Cochlea.
Vestibular
membrane
Tectorial
membrane
Basilar
membrane
From oval
window
To round
window
Diagrammatic and sectional views of the cochlear spiral
b
Temporal bone
(petrous part)
Scala
vestibuli
(contains perilymph)
Cochlear duct
(contains
endolymph
)
Spiral organ
Spiral ganglion
Scala
tympani
(contains perilymph)
Cochlear nerve
Vestibulocochlear
nerve (VIII)Slide51
Figure 17-28a The Spiral Organ.
Bony cochlear wall
Scala
vestibuli
Vestibular membrane
Cochlear duct
Tectorial membrane
Basilar membrane
Scala
tympani
Spiral organ
Spiral
ganglion
Cochlear branch
of N VIII
A three-dimensional section of the
cochlea, showing the compartments,
tectorial membrane, and spiral organ
aSlide52
Figure 17-28b The Spiral Organ (Part 1 of 2).
Tectorial membrane
Outer
hair cell
Basilar
membrane
Inner
hair cell
Nerve
fibers
Diagrammatic and sectional views of the receptor
hair cell complex of the spiral organ
bSlide53
Figure 17-28b The Spiral Organ (Part 2 of 2).
Cochlear duct
Vestibular membrane
Tectorial membrane
Scala
tympani
Basilar
membrane
Hair cells
of spiral
organ
Spiral ganglion
cells of
cochlear nerve
Spiral organ
LM
×
125
Diagrammatic and sectional views of the receptor
hair cell complex of the spiral organ
bSlide54
17-5 The EarAn Introduction to Sound
Pressure waves Consist of regions where air molecules are crowded together Adjacent zone where molecules are farther apart Sine wavesS-shaped curvesSlide55
17-5 The EarPressure Wave
WavelengthDistance between two adjacent wave troughsFrequencyNumber of waves that pass fixed reference point at given time Physicists use term cycles instead of wavesHertz (Hz) number of cycles per second (cps)Slide56
17-5 The EarPressure Wave
Pitch Our sensory response to frequencyAmplitudeIntensity of sound wave Sound energy is reported in decibelsSlide57
Figure 17-29a The Nature of Sound.
Wavelength
Tympanic
membrane
Air
molecules
Tuning
fork
Sound waves (here, generated by a tuning fork)
travel through the air as pressure waves.
aSlide58
Figure 17-30 Sound and Hearing (Part 1 of 2).
External
acoustic
meatus
Malleus
Incus
Stapes
Oval
window
Movement
of sound
waves
Tympanic
membrane
Round
window
1
2
3
Sound wavesarrive attympanicmembrane.
Movement ofthe tympanicmembrane causes
displacementof the auditoryossicles
Movement of
the stapes at
the oval window
establishes
pressure waves
in the perilymph
of the
scala
vestibuli
.
1
2
3Slide59
Figure 17-30 Sound and Hearing (Part 2 of 2).
5
4
6
4
5
6
Cochlear branch
of cranial nerve VIII
Scala
vestibuli
(contains perilymph)
Vestibular membrane
Cochlear duct
(contains
endolymph
)
Basilar membrane
Scala
tympani
(contains perilymph)
The pressure
waves distort
the basilar
membrane on
their way to the
round window
of the
scala
tympani.
Vibration of
the basilar
membrane
causes hair cells
to vibrate against
the tectorial
membrane.
Information about
the region and
the intensity of
stimulation is
relayed to the CNS
over the cochlear
branch of cranial
nerve VIII.Slide60Slide61
17-5 The EarAuditory Pathways
Cochlear branchFormed by afferent fibers of spiral ganglion neuronsEnters medulla oblongata Synapses at dorsal and ventral cochlear nuclei Information crosses to opposite side of brainAscends to inferior colliculus of midbrainSlide62
17-5 The EarAuditory Pathways
Ascending auditory sensationsSynapse in medial geniculate nucleus of thalamusProjection fibers deliver information to auditory cortex of temporal lobeSlide63
Figure 17-32 Pathways for Auditory Sensations (Part 1 of 2).
Primary pathway
Secondary pathway
Motor output
KEY
Stimulation of hair cells at
a specific location along
the basilar membrane
activates sensory neurons.
Sensory neurons carry the
sound information in the
cochlear branch of the
vestibulocochlear
nerve
(VIII) to the cochlear
nucleus on that side.
Cochlea
Low-frequencysounds
High-frequencysounds
Vestibularbranch
Vestibulocochlearnerve (VIII)
1
2Slide64
Figure 17-32 Pathways for Auditory Sensations (Part 2 of 2).
Primary pathway
Secondary pathway
Motor output
KEY
3
4
5
6
Projection fibers then deliver
the information to specific
locations within the auditory
cortex of the temporal lobe.
Low-frequency
sounds
Ascending acoustic
information goes to the
medial geniculate nucleus.
The inferior
colliculi
direct a variety of
unconscious motor responses to sounds.
To reticular formation
and motor nuclei of
cranial nerves
Superior
olivary
nucleus
Information ascends from each cochlear
nucleus to the superior
olivary
nucleus of the
pons and the inferior
colliculi
of the midbrain.
High-
frequency
sounds
Thalamus
To
ipsilateral
auditory
cortex
Motor output
to spinal cord
through the
tectospinal
tractsSlide65
17-5 The EarHearing Range
From softest to loudest represents trillionfold increase in powerNever use full potential Young children have greatest rangeSlide66
Table 17-1 Intensity of Representative Sounds.Slide67
17-5 The EarEffects of Aging on the Ear
With age, damage accumulatesTympanic membrane gets less flexibleArticulations between ossicles stiffenRound window may begin to ossify