1 THE EARTH THROUGH TIME TENTH EDITION H A R O L D L L E V I N Chapter 17 Human Origins 2013 JOHN WILEY amp SONS INC ALL RIGHTS RESERVED 2 Introduction Humans Homo sapiens meaning thinking man or intelligent human appeared during the latter part of Cenozoic during P ID: 525841
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Slide1
© 2013 JOHN WILEY & SONS, INC. ALL RIGHTS RESERVED.
1
THE EARTH THROUGH TIME
TENTH EDITION
H A R O L D L. L E V I NSlide2
Chapter 17
Human Origins
© 2013 JOHN WILEY & SONS, INC. ALL RIGHTS RESERVED.
2Slide3
Introduction
Humans, Homo sapiens, meaning "thinking man" or "intelligent human," appeared during the latter part of Cenozoic, during Pleistocene.Homo sapiens share many characteristics with the other members of the Order Primates, including basic body structure and biochemistry.
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3Slide4
Characteristics of Homo sapiens
Humans are distinct from the other primates : Larger, more complex brain Stand and walk erect due to structural modifications to vertebral column, legs & pelvis
Flatter face
Teeth less robust
Greater manual dexterity, leading to ability to manufacture and use sophisticated tools
Greater intelligence, leading to language and culture
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4Slide5
Order Primates
Primates are placental mammals. They have five digits, which is a primitive, non-specialized characteristic. They have not developed hoofs, horns, antlers, or trunks, unlike some other groups of placental mammals, and so they remain structurally generalized
compared with other mammalian groups.
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5Slide6
Adaptations of the Primates
Rounded head shapeProgressive enlargement of the brain
Shortening and flattening of the face
Unspecialized teeth
Eyes are close-set and positioned toward front of face.
Binocular stereoscopic vision
. Ability to judge distance.
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6Slide7
Adaptations of the Primates
Modifications of the hand - opposable thumb Mobility of the forearm, so hand can be turned
Modifications of the thorax for upright posture
Forelimbs and hind limbs diverged in form and function
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7Slide8
Right hand of the Eocene prosimian
Europolemus. The hand shows key primate characteristics: broad fingertips and an opposable thumb (the thumb can be touched to each of the other fingertips).
Right hand of a human (palm up).
The human hand is not used in locomotion. It can be used to manipulate small objects between the fingers and thumb.
The Opposable Thumb
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FIGURE 17-2
8Slide9
Types of Modern Primates
Prosimians - tree shrews, lemurs, lorises, and tarsiers Anthropoidea - monkeys, apes, and humans
Monkeys
New World monkeys
Old World monkeys
Hominoidea
or anthropoid apes
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9Slide10
Prosimians
Tree shrews, lemurs, lorises, and tarsiers Tree shrews and lemurs have long snouts and eyes on the sides of the head. Most digits have claws.
Tarsiers have relatively flat face, close-set eyes for stereoscopic vision, and fingers and toes have nails rather than claws.
lemurs
tarsier
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10Slide11
Not involved in evolution of humans
Includes spider monkey, capuchin, and marmoset.
Prehensile tails
Oldest fossils are Oligocene from South America
New World Monkeys
(
A) Art Wolfe/Photo Researchers, Inc.,
(B)
Jany
Sauvanet
/Photo Researchers, Inc
.,
(
C)
Gregory G
.
Dimjian
/Photo Researchers,
Inc
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11Slide12
Old World Monkeys
Baboons, mandrills, macaques, rhesus monkeys, Barbary apes
Nostrils close together and directed downward, as in humans
Tail not prehensile
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12Slide13
Macaque Baboons Barbary apes
Old World Monkeys
(left) Richard T.
Nowitz
/Photo Researchers, Inc.; (center) Tom McHugh/Photo Researchers, Inc.; (right) Len Rue, Jr./Photo Researchers, Inc.
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13Slide14
Hominoidea or Anthropoid Apes
Tail-less primates
Gibbons, orangutans, chimpanzees,
bonobos
, gorillas, and humans
Modern species evolved from same ancestral stock that produced humans
DNA evidence indicates divergence from human line 5-7 million years ago.
DNA of chimpanzees and humans is 98.4% similar.
Similarities in the proteins hemoglobin and
myoglobin
indicate that the chimpanzee is our closest relative.
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14Slide15
Hominoidea or Anthropoid Apes
Three families:Hylobatidae - Gibbons The most primitive branch of tail-less apes.
Pongidae
- Orangutans, chimpanzees, gorillas, and bonobos.
Hominidae
- Humans
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15Slide16
Anthropoid Apes
Gibbon
Gorilla
Orangutan
Chimpanzees
(top left) David Maitland/Getty Images, Inc.; (top right) Tom McHugh/Photo Researchers, Inc.; (bottom left) Andrew
Dernie
/Getty Images, Inc.; (bottom right)
Photodisc
/Getty Images, Inc.)
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16Slide17
Prosimian Vanguard
Earliest fossil record for primates is Purgatorius, known from the Upper Cretaceous Hell Creek Formation at Purgatory Hill, Montana. The earliest primates lived during the time of the last of the dinosaurs.
The
Paleocene Plesiadapis
is the only primate genus other than
Homo
to inhabit both the Old World and the New World. This suggests that the continents were not fully separated by Paleocene.
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17Slide18
Prosimian Vanguard
During Eocene, primates underwent: Reduction in length of face Increase in brain size Movement of eyes to more forward position Development of a grasping big toe
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18Slide19
Prosimian Vanguard
Tarsiers and lemurs were abundant and widely distributed on Northern Hemisphere continents during Eocene. When the climate cooled during Oligocene, the primates virtually disappeared from North America. They were forced southward into Asia, Africa, and the East Indies.
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19Slide20
Early Anthropoids
Anthropoids are the higher primates - monkeys, apes, and humans
Early anthropoid fossils are found in
Oligocene
rocks (33-34
m.y
. old) at Fayum, Egypt.
Aegyptopithecus
=
robust arboreal primates with a monkey-like tail and limbs, and eyes on the front of the skull.
The
prosimian
-anthropoid transition had occurred by Oligocene.
Oligocene ape, Aegyptopithecus
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20Slide21
Primate Evolution
During Miocene (23 - 5 m.y. ago), plate tectonics affected primate evolution. Africa and Arabia drifted northward and collided with Eurasia. This changed the circulation in the Tethys Sea and the climate in East Africa became cooler and drier.
Grass-covered plains replaced tropical forests.
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21Slide22
Primate Evolution
As a result of the change in climate and vegetation, adaptive radiation occurred among the Old World primates. During late Miocene, primates appeared that gave rise to the
pongids
(orangutan, chimpanzee, and gorilla) and the
hominids
(human family).
A new group, the
dryomorphs
(formerly dryopithecines) appeared during Miocene.
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22Slide23
The Dryomorphs - Proconsul
Proconsul, discovered by the Leakeys in 1948 in Kenya, was a
dryomorph
or a
dryomorph
ancestor.
It had ape-like skull, jaws, and teeth. It had monkey-like long trunk, arms and finger bones.
Middle Miocene
descendants of
Proconsul
are probably ancestral to modern African apes and the first hominids, the australopithecines.
Miocene
drymorph
,
Proconsul
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Harold Levin
23Slide24
Primate Evolution
By about 18 million years ago, Africa collided with Eurasia and primates migrated into Eurasia and diversified. These primates include the following Miocene apes, called
ramamorphs
, which are considered to be ancestral stock for both later apes and hominids:
Ramapithecus
Sivapithecus
Gigantipithecus
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24Slide25
Primate Evolution
Interpreting the primate family tree is complicated. There are many splits and dead ends. The story must be reinterpreted each time new types of fossil primates are discovered.
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25Slide26
Rich Fossil Sites of Africa
Rich hominin fossil sites in East Africa. Beds of volcanic ash between fossil beds facilitate radioisotope dating in this region, and dry conditions limit the amount of vegetation covering bone beds.
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26Slide27
Australopithicines
Australopithecus africanus was discovered in 1924 by Raymond Dart in South Africa. Since then, many specimens have been discovered in East Africa, particularly in Olduvai Gorge
by
Mary and Louis Leakey.
East African fossil sites have yielded hundreds of hominid bones, documenting human evolution over the past 4 million years.
Interbedded
volcanic ash
allows
radiometric dating
of the hominid fossils using potasium-argon method.
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27Slide28
Australopithecus africanus
Australopithecus africanus
skull, from the Transvaal region, South Africa
.
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28Slide29
Oldest hominid fossils are 6-7 million years old, from Chad, Sahelanthropus tchadensis.
Prior to this fairly recent discovery, the oldest know hominids were 5.8-5.2 million years old. Ardipithecus ramidus from Ethopia. Australopithecus anamensis
lived from 4.2 to 3.9 m.y. ago. It appears to be an evolutionary intermediate between
Ardipithecus ramidus
and
Australopithecus afarensis
or "Lucy."
Australopithicines
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29Slide30
"Lucy" Australopithecus afarensis
Discovered by Donald Johanson in 1974 and nicknamed "Lucy."
Lucy was a 3.5 foot tall
erect-walking hominid
in East Africa about 3.5 million years ago. Posture is determined from analysis of pelvic and leg bones.
Resent excavations for
Australopithecus
afarensis
have yielded a skeleton of a male that was about 5.5 feet tall.
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30Slide31
"Lucy" Australopithecus afarensis
Further evidence of bipedalism (walking upright on two feet) of australopithecines is the discovery of
footprints and a trackway
in 3.2 m.y. volcanic ash layers (a Pliocene tuff) at Laetoli, eastern Africa.
Australopithicine
dentition
(tooth structure) was
essentially human. Cranial (brain) capacity was less
than that of modern humans. (600 cm³ vs. 1400-1600³ in modern humans).
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31Slide32
Australopithecus afarensis
Evidence for bipedalism. (A) Footprints probably made by Australopithecus
afarensis
about 3.5 million years ago as a pair of these
hominins
walked across wet volcanic ash. The footprints document the ability of hominins to walk on two legs. The tracks were discovered in 1976 by Mary Leakey. (B) A female and male stroll the Late Pliocene landscape of eastern Africa, leaving telltale footprints in the ash, much as vacationers leave footprints on a wet beach.
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32Slide33
Two types of Australopithicines
Gracile - Smaller, lighter-bodied with smaller teeth. Includes Australopithecus afarensis (Lucy).
This group is transitional to genus Homo.
Robust
- Larger, heavy-bodied with larger teeth.
Includes
Australopithecus boisei
(sometimes called
Paranthropus boisei
) and
Australopithecus robustus
(sometimes called
Paranthropus robustus
). Both are evolutionary side branches.
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33Slide34
Opinions differ as to whether the boisei and robustus
species should be included in genus Australopithecus. The current consensus in the scientific community is that they should be placed in a separate genus, Paranthropus, which appears to have developed from the australopithecines.
The robust australopithecines are now reclassified as belonging to genus
Paranthropus
.
Two types of
Australopithicines
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34Slide35
New discover near Johannesburg, South Africa named Australopithecus sediba
Skeletal remains of the young male and an adult female.Intermediate between australopithecine and genus Homo. Australopithecine traits – small brain size, shoulder bones, and long arms.
Homo traits – modern hands and less flared pelvis
Transition between
Australopithicines
and
Homo
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35Slide36
Timelines for Pliocene and Pleistocene
hominins.
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36Slide37
Genus Homo
The genus Homo arose nearly 2.5 million years ago when australopithecines evolved into the ancestors of humans, Homo ergaster, also called Homo habilis
.
Homo rudolfensis
and
Homo ergaster
or Homo habilis lived in Africa around 2 million years ago.
Homo rudolfensis
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37Slide38
Genus Homo
The evolutionary transition may have been stimulated by the change to a cooler, drier climate about 2.7 million years ago.
Rainforests were replaced by grasslands.
In this environment, selective pressures may have led to
bipedalism
, greater
intelligence
, and the ability to make
stone tools
. Stone tools are found with fossils of
Homo
.
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38Slide39
Genus Homo
Genus Homo has a larger cranial capacity and some smaller teeth, but no striking anatomical differences.
Homo erectus
arose from
Homo ergaster or Homo habilis
.
The opening at the base of the skull, where the spinal cord joins the brain, called the foramen magnum, is in a more forward position in
Homo erectus
, indicating that it had a more erect posture.
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39Slide40
The Homo erectus Stage
Homo erectus fossils include: A 1.8 million years jaw in the Caucasus Mountains of Georgia
A 1.5 million years complete skeleton from east Africa near Lake Turkana (long-limbed, tall and slender with a remarkably modern skeleton)
A 750,000 year old skull cap from Olduvai Gorge
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40Slide41
Homo erectus lived in the early to middle Pleistocene
Homo erectus is the first hominid known to have moved from Africa into Eurasia. A rapid increase in brain size had begun in Homo erectus. Homo erectus
was a
toolmaker
and
hunter
. It is unclear whether they had language, wore clothes, built dwellings or used fire.
The Homo erectus Stage
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41Slide42
Homo erectus skull characteristics
Cranial capacity was 775 - 1300 cm³, slightly less than (or overlapping the lower range of) that of modern humans. Skull was rather massive and flat Heavy, bony supraorbital (brow) ridges
Sloping forehead
The jaw jutted forward at the toothline, a condition called
prognathous
The nose was broad and flat
The teeth were robust but essentially modern
Lacked a jutting chin
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42Slide43
Homo erectus and Tools
Progressive improvement in tool-making from stone during the Pleistocene. The crude stone tools of the Early Pleistocene (bottom) were produced by australopithecines. Homo erectus produced the better-shaped tools of the Middle Pleistocene. The Upper Paleolithic tools included carefully chipped blades and points.
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FIGURE
17-26, Harold Levin
43Slide44
Homo erectus and Fire
A group of Homo erectus people using fire to make tools while others return from hunting.
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44Slide45
The Neandertals
Late Pleistocene hominids are called Neandertals.
Neandertals
may be a subspecies
of
Homo sapiens
, designated as Homo sapiens neandertalensis.
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45Slide46
Neandertal Skull Characteristics
Brain size equal to or larger than that of modern humans Heavy supraorbital ridge
(brow ridge)
Prognathous
(face extends forward at toothline) - but not as much as
Homo erectus
. No chin - but lower jaw not as receding as that of Homo erectus
Enlarged nasal cavity (for warming cold inhaled air)
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46Slide47
Neandertal Skull Characteristics
Skull of a classic Neandertal.
From the
Homo erectus
stage of the Middle Pleistocene.
Neandertals
roamed central Europe as recently as 28,000 years ago.
This is indicated by the radiocarbon date for this
Neandertal
jawbone from a cave site in Croatia.
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Harold Levin
47Slide48
Neandertal Characteristics
Neandertal skeleton (A) assembled from actual skeletal bones compared with skeleton of a modern human. (B) The
Neandertal’s
flaring rib cage provided space for large lungs, needed for a high level of activity.
Reconstruction of a Neandertal family group.
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48Slide49
When Did the Neandertals
Live?The Neandertals can be divided into three groups. Early Neanderthals, that lived approximately 250,000 to 130,000 years ago
Neanderthals that existed during the transition to the Upper Paleolithic, approximately 130,000 to 45,000 years ago
Late-surviving Neanderthals after 45,000 years ago, up until about 28,000 years ago.
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49Slide50
Neandertals and Modern Humans
In central Europe, Neandertals coexisted with early modern humans for thousands of years.
Neandertal skeletons were somewhat more robust than human skeletons.
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50Slide51
Neandertals and Modern Humans
Bones discovered in 2002 show a mix of human and Neandertal traits, suggesting interbreeding.
Skull comparison.
(A)
Neandertal
. (B) Skull from a rock shelter on the slope of Mount Carmel (Israel) that appears to show both
Neandertal
and Cro-Magnon features. (C) Cro-Magnon. The Mount Carmel skull is intermediate in both form and age between
Neandertal
and Cro-Magnon.
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Harold Levin
51Slide52
Neandertal Culture
Neandertals hunted cold-tolerant mammals including cave bears, mammoths, wooly rhinos, reindeer, bison, and aurochs (a type of cattle).
They manufactured a variety of
stone tools
They used
fire
, which provided
light
in caves, gave
warmth
, allowed the
thawing and cooking
of food, and provided
protection from predators.
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52Slide53
Neandertal Culture
They constructed shelters or houses of skins, sticks and bones.
They
cared for the sick
.
They
buried artifacts with the dead, suggesting a belief in an afterlife.
Musical instruments
have also been found in association with Neandertal remains, such as a flute made from a bone, dating to between 82,000 and 43,000 years ago.
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53Slide54
Little People of the South Pacific
A new species of human was discovered on the Indonesian island of Flores in 2004, Homo floresiensis. An excavation produced the remains of seven small individuals.
The adults were only 3 ft (1 m) tall.
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54Slide55
Little People of the South Pacific
Homo floresiensis lived as recently as 13,000 years ago, at the same time as both the Neandertals and modern humans. They hunted, used fire, and made stone tools.
The ancestry of this species is uncertain, however,
Homo erectus
probably reached the island about 800,000 years ago, and evolved into a smaller species as a result of living on an island with limited food and few large predators.
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55Slide56
The Earliest Humans
Fossil evidence from along the Omo River in southern Ethiopia shows that modern humans, Homo sapiens, lived in Africa as much as 195,000 years ago, according to an article published in February 2005
Previously, modern humans were thought to have appeared in Africa about 160,000 years ago
Humans appeared in Africa many thousands of years before our species appeared on any other continent
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56Slide57
The Earliest Humans
There appears to have been a time gap between the appearance of the modern human skeleton and modern types of behavior resulting in cultural artifacts.
Stone knife blades appeared between 200,000 and 50,000 years ago.
Other types of cultural artifacts, however, appeared just 50,000 years ago.
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57Slide58
Out of Africa
Humans left Africa between about 40,000 and 50,000 years ago, and appear to have taken culture with them, including things such as bone carving, tools such as harpoons, jewelry and ornamentation, artwork, and arrowheads.
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58Slide59
Cro-Magnon
About 34,000 years ago during a glacial advance, humans closely resembling modern Europeans moved into the area occupied by the Neandertals, and eventually replaced them. They are Homo sapiens, and are called
Cro-Magnon.
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59Slide60
Cro-Magnon
Characteristics of Cro-Magnon: Mostly taller than Neandertals More vertical forehead Jutting chin
Cro-Magnon skull, 30,000 years old
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60Slide61
Cro-Magnon Culture
Cro-Magnon continued and further developed the cultural traditions of the Neandertals:Manufactured a greater variety of stone tools
Painted pictures of animals in caves
Carved and sculpted
images of women and animals from bone or ivory
Made and wore
jewelry
Elaborately buried their dead (such as burying hunters with their weapons)
Practiced rituals and believed in afterlife
Hunted and gathered edible plants
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61Slide62
Cro-Magnon Culture
Prehistoric art by Late Pleistocene Homo sapiens. (A) “The Venus of Willendorf”—probably a fertility figure—found in Austria. (B) A tool called a thong-
stropper
, used either to work thongs made from hide or to straighten arrow shafts. The tool is made from an antler and is intricately carved.
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62Slide63
Beginnings of Recorded History
About 15,000 -10,000 years ago, humans began to: Domesticate animals
Cultivate plants
Produce more highly developed tools
Make
utensils of fired clay
Develop permanent settlements
Use
language
Develop
writing
With the development of writing, the era of recorded history began.
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63Slide64
Coming to America
Humans in the Americas are called Paleoindians. It is not definitely known when the first human set foot in America.
The widely held view is that humans arrived in the Americas from Asia, crossing the
Bering Land Bridge
between Asia and Alaska between 10,000 and 25,000 years ago (to perhaps 30,000 years or more), during a glaciation which lowered sea levels.
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64Slide65
Pre-Clovis People
Stone weapon points and tools in Meadowcroft Shelter, western Pennsylvania, appear to be 19,600 years old.Some South American sites are 13,000 years old or older.
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65Slide66
Arrival of the Clovis People
Stone tools of the Clovis culture, about 13,000 years old, are known from Alaska to South America. They occupied sites in the Americas for over 2,000 years.
The Clovis people dominated for several thousand years, and then a new group, the
Folsom people
took up residence in North America about 11,000 years ago.
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66Slide67
Human Population – 7 billion and growing
Human survival may depend on our ability to slow the rate of population increase.
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67Slide68
What lies ahead?
Humans have existed for about the past 2 million years out of the Earth’s 4.6 billion year history. Will humans match the Dinosaurs lineage of 140 million years?Global climates and sea level will be influenced by both human (green house gases) and natural (
Milankovitch
cycles) processes.
The Earth is still tectonically active with the Atlantic ocean basin continuing to widen, Africa moving north to collide with Europe, and the Baja Peninsula on an Alaskan cruise.
In 7 billion years the Sun will expand as part of the red giant phase and engulf the Earth.
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68Slide69
FIGURE 17-2 Right hand of a human (palm up). Source: The Primate Order Table, Copyright © 2000-2012 by Dennis O'Neil, located at http://anthro.palomar.edu/primate/table_primates.htm
IMAGE CREDITS
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69