1 THE EARTH THROUGH TIME TENTH EDITION H A R O L D L L E V I N Chapter 12 Life of the Paleozoic 2013 JOHN WILEY amp SONS INC ALL RIGHTS RESERVED 2 Paleozoic overview FIGURE 101 Major events of the Paleozoic Era ID: 755755
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Slide1
© 2013 JOHN WILEY & SONS, INC. ALL RIGHTS RESERVED.
1
THE EARTH THROUGH TIME
TENTH EDITION
H A R O L D L. L E V I NSlide2
Chapter 12Life of the Paleozoic
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2Slide3
Paleozoic overview
FIGURE 10-1 Major events of the Paleozoic Era.
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3Slide4
Paleozoic Fossil Record In Paleozoic rocks, we find abundant fossils of multicellular organisms bearing shells.
The fossil record is much improved at the beginning (the base of) Paleozoic strata. The pace of evolution appears to have quickened in the Paleozoic
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4Slide5
Paleozoic InvertebratesRepresentatives of most major invertebrate phyla were present during Paleozoic, including sponges, corals, bryozoans, brachiopods, molluscs, arthropods, and echinoderms.
Almost all of the common invertebrate phyla in existence today had appeared by Ordovician.
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5Slide6
Paleozoic VertebratesVertebrates evolved during Paleozoic, including:
FishesAmphibiansReptilesSynapsids ("mammal-like reptiles")
The first vertebrates were
jawless fishes
, which are found in rocks as old as Cambrian in China.
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6Slide7
Paleozoic VertebratesAn advanced lineage of fishes
with primitive lungs and stout fins gave rise to the four-legged animals or tetrapods.The transition from water-dwelling vertebrates to land-dwelling vertebrates depended on the evolution of the
amniotic egg
.
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7Slide8
Paleozoic PlantsThe first primitive land plants
appeared near the end of Ordovician. Vascular plants expanded across the land, forming great forests during Devonian. The plants progressed from seedless, spore-bearing plants to plants with seeds but no flowers (gymnosperms).
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8Slide9
Paleozoic ExtinctionsSeveral mass extinctions
occurred during Paleozoic, including the largest extinction of all at the end of Permian. Other mass extinctions occurred at the end of Ordovician and
Devonian
.
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9Slide10
Paleozoic Life Summary of invertebrate phyla
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10Slide11
Paleozoic Life cont.
Summary of invertebrate phyla
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11Slide12
Adaptive Radiations and Extinctions
Paleozoic was a time of several adaptive radiations and extinctions. Many geologic periods began with adaptive radiations (times of rapid evolution).
Several
periods ended with extinction events
of varying severity.
The extinction event at the end of
Permian
was the
most extensive mass extinction in the history of life.
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12Slide13
Diversity during Paleozoic Red arrows mark extinction events.
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FIGURE
12-94
13Slide14
Soft-bodied AnimalsMulticellular animals evolved during Precambrian.
Soft-bodied Ediacaran-type organisms ranged into Cambrian. Soft-bodied fossils are infrequently preserved.
Preservation improved with the origin of hard parts
.
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14Slide15
The first animals with shells are called small shelly fossils.Small shelly fossils are found
at the base of Cambrian, and during Late Neoproterozoic. Most disappeared during the Early Cambrian.
Small Shelly Fossils
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FIGURE
12-2
15Slide16
Small Shelly FossilsMany had phosphatic
shells, few mm in size. Shells and skeletal remains of primitive molluscs, sponges, and animals of uncertain classification, such as Cloudina, that secreted a calcareous tube.
FIGURE
12-1 Tiny
shell-bearing fossils from the Late
Precambrian and Early Cambrian in Siberia.
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16Slide17
Cambrian DiversificationThe initial Paleozoic diversification is known as "the Cambrian explosion
." Abrupt appearance of many types of animals about 535 million years ago, followed by rapid evolution.During that episode of explosive evolution, all major invertebrate phyla appeared in the fossil record (except Bryozoa).
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17Slide18
Cambrian Substrate RevolutionInfaunal, burrowing animals evolved rapidly during Cambrian, as indicated by
trace fossils and bioturbation (disruption of sedimentary structures by biological activity) of sediments. The dramatic change in the character of the seafloor sediments (from undisturbed to highly burrowed) has been called the "
Cambrian substrate revolution
.
"
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18Slide19
Why the Cambrian Explosion?No satisfying answers to why life diversified at the beginning of the Cambrian. The answer likely involves a number of factors.
Climate conditions became more favorable after the end of the Neoproterzoic glaciation.Perhaps the glaciation produced an extinction event in the Ediacaran animals.Extinction events of the Phanerozoic have been followed by rapid adaptive radiation
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19Slide20
Soft-Bodied Fossils in the Burgess Shale
The extraordinarily well-preserved Middle Cambrian Burgess Shale fauna of Canada provides a window into the past to view the spectacular diversity of Middle Cambrian. Many soft-bodied organisms
are preserved in black shale, along with the
soft parts of animals with shells
, such as legs and gills of trilobites.
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20Slide21
Soft-Bodied Fossils in the Burgess Shale The significance of the Burgess Shale is that is records
soft-bodied organisms, and the soft parts of organisms with shells. The finely detailed preservation reveals the extraordinary diversity and evolutionary complexity that existed near the beginning of Paleozoic.
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21Slide22
Stratigraphic setting of the Cambrian Burgess Shale
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FIGURE 12-5 The Burgess Shale fauna.
22Slide23
Animals in the Burgess Shale Several groups of arthropods, including trilobites and crustaceans
Sponges Onycophorans Crinoids
Molluscs
Corals
Three phyla of worms
Chordates (
Pikaia
)
Many others
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23Slide24
Location of the Burgess Shale fauna in British Columbia, CanadaC = Onycophoran, Aysheaia, intermediate in evolution between segmented worms and arthropods.D = Arthropod Leanchoila
E= Arthropod Waptia
Animals in the Burgess Shale
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FIGURE 12-5 The Burgess Shale fauna.
24Slide25
Animals in the Burgess Shale: Chordates
Chordates have a notochord or dorsal stiffening rod associated with a nerve chord, at some stage in their development. In vertebrates, the notochord is surrounded by and usually replaced by a vertebral column during embryonic development.
Vertebrates are chordates
, but
Pikaia
pre-dates the evolution of vertebrae.
It is thought that vertebrates evolved from organisms similar to Pikaia.
Pikaia
is a fish-like lower chordate from the Burgess Shale.
Modern representatives are called lancelets, such as the genus
Amphioxus
.
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FIGURE
12-13
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Predators in the Cambrian Seas
The giant predator of the Cambrian seas, Anomalocaris, up to 60 cm long. Predators would have caused selective pressures on prey. The need to avoid being eaten probably encouraged the evolution of hard protective shells.
Predation probably also caused an
increase in diversity
of prey, as they evolved to better survive predation.
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FIGURE 12-8
Anomalocaris
, “invertebrate equivalent of
the dinosaurs.”
26Slide27
Other Burgess Shale AnimalsMarrella,
a "lace crab," is common in the Burgess Shale. Hallucigenia, an onycophoran, was originally interpreted to walk on its spines, until claws were discovered on its "tentacles."
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FIGURE 12-10
FIGURE
12-11
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Exceptional PreservationFossil sites containing abundant fossils with extraordinary preservation are called
lagerstätten. Both the Burgess Shale fauna and the Chengjiang fauna from China are considered to be lagerstätten.
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28Slide29
The Chengjiang fauna In 1984, the Lower Cambrian Chengjiang fossil site was discovered in Yunnan Province, China.
More than 100 species of invertebrates have been found, with extraordinary preservation, including many soft bodied forms.
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29Slide30
The Chengjiang fauna Jelly fish
Annelid wormsCnidaria Porifera (sponges) Brachiopods ArthropodsEarly chordates similar to Pikaia
The world's oldest known fish
(
Myllokunmingia
)
Other species of unknown phyla
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30Slide31
Oldest Known Fish
The world's oldest known fish, Myllokunmingia, from the Maotianshan Shale near Chengjiang, in the Yunnan Province of China. 535 million years old.
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31Slide32
Ordovician Diversity
Following a slight dip in diversity at the end of Cambrian, Ordovician seas experienced renewed diversification.Global diversity tripled over a 25 million year time interval.
The number of genera increased rapidly, and the number of families increased from about 160 to 530.
The increase was particularly notable among
trilobites, brachiopods, bivalve
molluscs
, gastropods, and corals.
Why diversify?
Fragmented continents
Extensive seafloor spreading
Extensive warm nutrient rich seas fostering plankton growth resulting in an expansion of the base of the food chain.
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32Slide33
Late Ordovician ExtinctionAn extinction event at the end of Ordovician
led to an abrupt decline in diversity. This extinction event was apparently related to the growth of glaciers in Gondwana, coupled with a reduction in shallow water habitat associated with the lowering of sea level.
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33Slide34
Diversity and extinction during Paleozoic Red arrows mark extinction events
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34Slide35
Silurian Diversity Diversification of marine animals occurred again at the beginning of Silurian.
The period ended with only a slight drop in diversity.
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35Slide36
Devonian Diversity During Devonian, there was continued diversification, but this
ended with another fairly large extinction event, which extended over about 20 million years. Roughly 70% of marine invertebrates disappeared. Because of the long duration, the extinction is unlikely to have been caused by a sudden, catastrophic event.
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36Slide37
Carboniferous-Permian Diversity During Early Carboniferous, diversity once again increased.
Diversity of marine animals remained fairly constant throughout Carboniferous and Permian. Late Permian is marked by a
catastrophic
extinction event which resulted in the total disappearance of many animal groups
.
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37Slide38
Overview of Changes in Diversity Through Time
Several Paleozoic periods ended with extinction events The beginning of most Paleozoic periods were marked by
adaptive radiations
Maximum diversity in Paleozoic seas was maintained roughly constant at between 1000 and 1500 genera
The largest extinction occurred at the end of Permian
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38Slide39
Recovery of diversity during Mesozoic was slow Diversity increased rapidly during Cretaceous Another mass extinction occurred at the end of Cretaceous
Diversity increased extremely rapidly, at unprecedented rates, at the beginning of Cenozoic Diversity during Cretaceous and Cenozoic was much greater than during Paleozoic
Overview of Changes in
Diversity Through Time
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39Slide40
Red arrows mark extinction events
Overview of Changes in
Diversity Through Time
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40Slide41
Unicellular Organisms in the Paleozoic Seas The principal groups of Paleozoic unicellular animals with a significant fossils record are the
foraminifera and the radiolaria, which belong to Phylum Sarcodina.
These organisms are
unicellular eukaryotic organisms
, and belong to
Kingdom Protista
.
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41Slide42
Foraminifera Name: Foraminifera means "hole bearer."
Chief characteristics: Unicellular. Related to the amoeba, with pseudopods. Foraminifera build tiny shells (called tests) which grow by adding chambers.
Some species (called agglutinated foraminifera) construct tests of tiny particles of
sediement
. This is the most primitive test.
Other
forams construct tests of calcium carbonate.
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42Slide43
Foraminifera
Geologic range: Cambrian to Holocene.Modes of life: Benthic or benthonic (bottom dwellers) Planktic or planktonic (floaters).
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43Slide44
Fusulinid foraminifera (fusulinids)
Fusulinids were abundant during Late Paleozoic (primarily Pennsylvanian and Permian).Their tests were similar in size and shape to a grain of rice. Their internal structure is complex and used to distinguish different species.Important guide fossils during Pennsylvanian and Permian because they
evolved rapidly
, were
abundant
, and
widespread geographically.
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44Slide45
Radiolaria
Chief characteristics: Unicellular. Test or shell composed of opaline silicaOrnate lattice-like skeleton
Often spherical or radially symmetrical with spines
Geologic range
:
Precambrian or Cambrian to Holocene. Rare during Early Paleozoic. More abundant during Mesozoic and Cenozoic
.
Mode of life
:
Planktonic. Marine only
.
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45Slide46
Radiolaria and the Rock RecordRadiolarians are
important constituents of chert at certain times in geologic history.Their tests accumulate on the seafloor today to form radiolarian ooze, particularly in deep water, where any calcium carbonate shells would be dissolved.
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46Slide47
Marine Invertebrates in the Paleozoic Seas The fossils of shell-bearing invertebrates that inhabited shallow seas are common in Paleozoic rocks.
Archaeocyathids, sponges, corals, bryozoans, trilobites, molluscs, and echinoderms. Many were benthic (bottom dwellers), but others, such as graptolites, were
planktonic
. Currents carried them over wide areas.
As a result, they are useful
index fossils for global stratigraphic correlation
.
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47Slide48
Phylum Archaeocyatha Name means "ancient cup"
Chief characteristics: Conical or vase-shaped skeletons made of calcium carbonate. Double-walled structure with partitions and pores. Geologic range: Cambrian only. Extinct. Mode of life: Attached to the sea floor. Reef-builders.
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FIGURE 12-17
Archaeocyathid
skeleton.
48Slide49
Phylum Porifera - The Sponges
Name means “pore-bearing,” or covered by tiny pores.
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FIGURE 12-21 Schematic diagram of a sponge with
the simplest
type of canal system.
FIGURE 12-18 Early Paleozoic sponges.
49Slide50
Phylum Porifera - The Sponges
Chief characteristics: Globular, cylindrical, conical or irregular shape. Basic structure is vase-like with pores and canals. Interior may be hollow or filled with branching canals.
Solitary or colonial.
Skeletal elements are called
spicules
, and they may be separate or joined.
Composition may be
calcareous, siliceous
or organic material called
spongin
.
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50Slide51
Phylum CnidariaCorals, sea fans, jellyfish, and sea anemones.
Name: Cnidaria are named for stinging cells called cnidoblasts or cnidocytes.Many are soft-bodied but only those which form hard skeletal structures are readily preservable as fossils.
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FIGURE 12-23 Common cnidarians.
51Slide52
Phylum CnidariaG
eologic range: Late Precambrian (Proterozoic) to Holocene for the phylum.The first corals were the tabulates. Mode of life: Corals live attached to the sea floor, primarily in warm, shallow marine environments.
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52Slide53
Phylum Cnidaria – Chief Characteristics
Radial symmetryMouth at the center of a ring of tentacles.
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FIGURE 12-24
53Slide54
Phylum Cnidaria – Chief Characteristics
Body form may be polyp (attached to the bottom, with tentacles on top) or medusa (free-swimming, jellyfish).
Diorama photograph courtesy of the U.S. National Museum of Natural History/
Smithsonian Institution.)
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54Slide55
Chief Characteristics of CoralsMay be
solitary or colonial. Colonies are composed of many polyps living together. Hard calcareous skeleton. The skeletal parts formed by polyps are called corallites
.
The "cup," in which an individual coral polyp sits, is called the
theca
. Each theca is small, and roughly circular or hexagonal.
The theca is divided internally by vertical partitions called
septae
, arranged in a radial pattern.
Harold Levin
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55Slide56
Chief Characteristics of CoralsTypes of corals are distinguished by presence or absence, and number of septae:
Rugose corals (or tetracorals) have septae arranged in multiples of four. Tabulate corals lack septae. Mesozoic and Cenozoic
scleractinian corals
(or hexacorals) have septae arranged in multiples of six.
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56Slide57
Rugose
Corals
Most rugose corals are solitary and conical (shaped like ice cream cones).
Septae are visible in the circular opening of the cone.
Some rugose corals are colonial, having hexagonal corallites with septae (such as
Hexagonaria
from Devonian of Michigan).
Harold Levin
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57Slide58
Rugose CoralsGeologic range: Ordovician to Permian - all extinct.
Rugose corals were abundant during Devonian and Carboniferous, but became extinct during Late Permian.
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58Slide59
Tabulate CoralsTabulate corals are
colonial and resemble honeycombs or wasp nests. They lack septae.They have horizontal plates within the theca called tabulae
. Tabulae are one of the main features of the tabulate corals.
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Harold Levin
59Slide60
Tabulate CoralsGeologic range: Ordovician to Permian - all extinct.
The principal Silurian reef formers.They declined after Silurian and their reef-building role was assumed by the rugose corals.
Harold Levin
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60Slide61
Modern CoralsModern corals are scleractinian corals. Scleractinian corals have septae are arranged in multiples of six, and are sometimes called
hexacorals. Scleractinian corals did not appear until after Paleozoic Geologic range: Triassic to Holocene.
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61Slide62
Phylum BryozoaName
: Name means "moss" (bryo) + "animal" (zoa). Chief characteristics: Colonial (many microscopic individuals living physically united adjacent to one another).The individuals are called zooids, and they are housed in a hard "capsule" called a
zooecium
.
The colony is called a
zoarium
.
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62Slide63
Phylum Bryozoa Individual
zooecia (plural of zooecium) are very tiny (about the size of a pin-hole, a millimeter or less in diameter). They are just large enough to be seen with the unaided eye. Bryozoans may be distinguished from corals because of the apertures in the skeleton are much smaller.
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63Slide64
Phylum Bryozoa The bryozoan colony may resemble lace or a tiny net, may be delicately branching, finger-like, circular or dome-shaped. There are more than 4000 living species of bryozoans, and nearly 16,000 fossil species.
Archimedes,
from
Mississippian rocks
, has a
cork-screw-like central axis
with a fragile net-like colony around the outer edge.
Harold Levin
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64Slide65
Phylum BryozoaGeologic range
: Ordovician to Holocene. Mode of life: Widespread in marine environments. A few live in freshwater lakes and streams.
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65Slide66
Phylum BrachiopodaName:
Name means "arm" (brachio) + "foot" (pod). Chief characteristics: Bivalved (two shells), each with bilateral symmetry. The plane of symmetry passes through the center of each shell or valve.
The two valves differ in size and shape
in most. Sometimes the larger valve will have an opening near the hinge line through which the
pedicle
extended in life.
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66Slide67
Phylum Brachiopoda Soft parts include a
lophophore consisting of coiled tentacles with cilia. The lophophore circulates water between the two valves, distributing oxygen and flushing out carbon dioxide. Water movements caused by the lophophore also transport food particles toward the mouth.
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67Slide68
Phylum Brachiopoda
Mode of life: Shallow marine environments. Generally attached to the sea floor.
Inarticulate brachiopods are known to live in burrows in the sediment.
Brachiopods are
filter feeders
.
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FIGURE 12-28
68Slide69
Inarticulate Brachiopods Primitive brachiopods with phosphatic or chitinous valves
.No hinge. Spoon-shaped valves held together with muscles and soft parts.Lingula is a well known genusGeologic range: Early Cambrian to Holocene
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69Slide70
Articulate Brachiopods Calcareous valves attached together with a hinge consisting of teeth and sockets.
Geologic range: Early Cambrian to HoloceneSpiny brachiopods (called productids) are characteristic of Carboniferous and Permian.
Harold Levin
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70Slide71
Phylum BrachiopodaGeologic range
: Early Cambrian to Holocene. Very abundant during Paleozoic. A few species (belonging to only three families) remain today.
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71Slide72
Phylum Mollusca Clams, oysters, snails, slugs,
Nautilus, squid, octopus, cuttlefish Name: Mollusca means " soft bodied."
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72Slide73
Phylum MolluscaChief characteristics
: Soft body enclosed within a calcium carbonate shell.A few, like slugs and the octopus, have no shell.Muscular part of body of clams and snails and some other groups of molluscs is called the foot.
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73Slide74
Phylum MolluscaGeologic range
: Cambrian to Holocene Mode of life: Marine, freshwater, or terrestrial. They may: swim, float or drift, burrow into mud or sand, bore into wood or rock, attach themselves to rocks, or crawl.
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74Slide75
Types of Molluscs
Monoplacophorans (Neopilina)Polyplacophorans or amphineurans (chitons)Bivalves or pelecypods (clams, scallops)Gastropods (snails and slugs)
Cephalopods (squid, octopus,
Nautilus
)
Scaphopods (tusk shells)
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75Slide76
Class Monoplacophora
Chief characteristics: Single shell resembling a flattened cone or cap. Soft part anatomy shows pseudo-segmented arrangement of gills, muscles, and other organs. Suggests that the primitive mollusc was a segmented animal. Segmentation was lost secondarily.
Monoplacophorans
are regarded as ancestral to bivalves, gastropods, and cephalopods.
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FIGURE 12-35
76Slide77
Class Monoplacophora Name
: Monoplacophora means "single plate-bearer."Geologic range: Cambrian-Holocene, but only known as fossils from Cambrian to Devonian. Living monoplacophorans found in deep water off Costa Rica in 1952 and named Neopilina. Considered to be a "living fossil."
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77Slide78
Class Amphineura or Polyplacophora – The Chitons
Chief characteristics: Chitons have 8 overlapping plates covering an ovoid, flattened body.
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FIGURE 12-34 A common
placophoran
,
the Atlantic Coast
chiton
.
78Slide79
Class Amphineura or Polyplacophora – The Chitons
Name: Polyplacophora means "many plate-bearer." Geologic range: Cambrian to Holocene
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79Slide80
Class Bivalvia or Pelecypoda
Clams, oysters, scallops, mussels, rudists Chief characteristics:Skeleton consists of two calcareous valves connected by a hinge.Bilateral symmetry; plane of symmetry passes between the two valves.
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FIGURE 12-36 Paleozoic bivalves
.
80Slide81
Class Bivalvia or Pelecypoda
Name: Bivalvia means " two" (bi) + " shells" (valvia). Geologic range: Early Cambrian to Holocene Mode of life: Marine and freshwater. Many species are infaunal burrowers or borers, and others are epifaunal.
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81Slide82
Class GastropodaSnails and slugs
Chief characteristics: Asymmetrical, spiral-coiled calcareous shell. Name: means "stomach" (gastro) + "foot" (pod).
Geologic range
: Early Cambrian to Holocene.
Mode of life
: Marine, freshwater or terrestrial.
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82Slide83
Class CephalopodaSquid, octopus,
Nautilus, cuttlefish Name: means " head" (kephale) + " foot" (pod). Chief characteristics: Symmetrical cone-shaped shell with internal partitions called septae
Shell may be
straight or coiled in a spiral which lies in a plane
.
Smooth or contorted
sutures visible on the outside of some fossils
mark the place where septae join the outer shell
.
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83Slide84
Class CephalopodaGeologic range
: Late Cambrian to Holocene Mode of life: Marine only; carnivorous (meat-eating) swimmers.
Types of Paleozoic cephalopods:
Nautiloids
Ammonoids
Coleoids
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84Slide85
Nautiloid CephalopodsThe shells of nautiloid cephalopods have smoothly curved septa, which produce simple, straight or curved sutures.
Geologic range: Cambrian to Holocene
Harold Levin
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85Slide86
Ammonoid CephalopodsAmmonoid cephalopods have complex, wrinkled or crenulated septa, which produce angular or dendritic sutures.
Geologic range: Devonian to Cretaceous - all extinct.
Harold Levin
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86Slide87
Ammonoid CephalopodsThere are three basic types of sutures in ammonoid shells:
Goniatite or goniatitic (septae have relatively simple, zig-zag undulations) Ceratite
or
ceratitic
(septae have smooth "hills" alternating with saw-toothed "valleys")
Ammonite
or
ammonitic
(septae are complexly branching and tree-like or dendritic)
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87Slide88
Types of Sutures in Cephalopods
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FIGURE 12-39 Cephalopod suture patterns.
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Subclass ColeoideaBelemnoids (belemnites)
Geologic range: Mississippian to Eocene - all extinct. Sepioids (cuttlefish)Geologic range: Jurassic to HoloceneTeuthoids (squid)
Geologic range
: Jurassic to Holocene
Octopods (octopus)
Geologic range
: Cretaceous to Holocene
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89Slide90
Order Belemnoidea - Belemnoids
The belemnoids have an internal calcareous shell (which resembles a cigar in size, shape, and color) called a rostrumThe front part of this shell is chambered, as in the nautiloids and ammonoids. The rostrum is made of fibrous calcite, arranged in concentric layers.
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90Slide91
Class ScaphopodaTusk shells or tooth shells
Chief characteristics: Curved tubular shells open at both ends. Geologic range: Ordovician to Holocene. Mode of life
: Marine.
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91Slide92
Phylum ArthropodaInsects, spiders, shrimp, crabs, lobsters, barnacles, ostracodes, trilobites, eurypterids
Name: means "jointed" (arthro) + "foot" (pod). Chief characteristics: Segmented body with a hard exterior skeleton composed of chitin (organic material). Paired, jointed legs.
Highly developed nervous system and sensory organs.
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92Slide93
Phylum ArthropodaGeologic range
: Cambrian to HoloceneMode of life: Arthropods inhabit a wide range of environments. Most fossil forms are found in marine or freshwater sediments.
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93Slide94
Paleozoic Arthropods and Their Geologic Ranges
Trilobites - Cambrian to PermianHorseshoe crabs - Silurian to HoloceneEurypterids - Ordovician to PermianArachnids - Late Silurian to HoloceneOstracodes - Cambrian to Holocene Onychophorans - Cambrian to HoloceneInsects - Devonian to Holocene
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Subphylum Trilobita - Trilobites
Chief characteristics: Body has three-lobes Skeleton composed of chitin, with calcium carbonate Body is divided into three segments:
Rigid head segment -
cephalon
Jointed, flexible middle section -
thorax
Rigid tail piece -
pygidium
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FIGURE 12-42
95Slide96
Subphylum Trilobita - TrilobitesName
: Trilobite means "three" (tri) + "lobed" (lobus). Geologic range: Cambrian to Permian Mode of life: Exclusively marine. Most were bottom dwellers living in shallow shelf environments.
Harold Levin
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96Slide97
Class
Eurypterida - EurypteridsExtinct scorpion-like or lobster-like arthropods.Predators. Up to 10 ft long.
Geologic range
: Ordovician to Permian. Most are Silurian or Devonian.
Mode of life:
Inhabited brackish estuaries.
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97Slide98
Class Arachnida - ArachnidsScorpions, spiders, ticks, and mites
Scorpions are the oldest arachnids with a fossil record. Scorpions had evolved by Late Silurian. The earliest ones appear to have lived in the water, because their fossils have gills. Scorpions, spiders, and mites are found in Devonian rocks. Geologic range: Late Silurian to Holocene.
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98Slide99
Class Ostracoda -Ostracodes
The ostracodes are mainly microscopic in size. Tiny bivalved shell encasing a shrimp-like creature. Geologic range: Cambrian to Holocene. Mode of life: Both marine and freshwater.
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99Slide100
Class OnychophoraOnychophorans share many characteristics of segmented annelid worms and arthropods, and are considered to be intermediate in evolution between the two groups.
Geologic range: Cambrian to HoloceneThe onycophoran,
Aysheaia
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100Slide101
Class Hexapoda - InsectsThe insects are among the most diverse living group on Earth, but they are rarely found as fossils.
Body is divided into three parts, head, thorax, and abdomen. Thorax has six legs. The earliest insects were wingless.Winged insects appeared by Pennsylvanian.
Geologic range
: Middle Devonian to Holocene.
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FIGURE
12-46
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Phylum Echinodermata Starfish, sea urchins, sand dollars, crinoids,
blastoids, and others Name: Echinodermata means "spiny" (echinos) + "skin" (derma).
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FIGURE 12-47 Representative living echinoderms.
102Slide103
Phylum Echinodermata
Chief characteristics: Calcite skeleton with five-part symmetry, superimposed on primitive bilateral symmetry. Echinoderms have a water vascular system
with water in a system of tubes within the body.
Tube feet
are soft, movable parts of the water vascular system which project from the body and are used in locomotion, feeding, respiration, and sensory perception.
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103Slide104
Phylum EchinodermataGeologic range
: Cambrian to Holcene. Mode of life: Exclusively marine. Some are attached to the sea floor by a stem with "roots" called holdfasts; others are free-moving bottom dwellers.Similarity of embryos between echinoderms and chordates suggests that they may be derived from a common ancestral form.
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104Slide105
Class Crinoidea - Crinoids
Crinoids are animals which resemble flowers.They consist of a calyx with arms, atop a stem of calcite disks called
columnals
.
The crinoid is attached to the sea floor by root-like
holdfasts
.
Some living crinoids are swimmers, and not attached.
Over 1000 genera are known.
FIGURE 12-55
Crinoid
in living position on
the seafloor
.
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105Slide106
CrinoidsGeologic range: Middle Cambrian to Holocene.
Especially abundant during Mississippian.
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Class Blastoidea - Blastoids
Blastoids are extinct animals with an armless bud-like calyx on a stem. About 95 genera are known. A common genus is Pentremites. Geologic range
: Ordovician to Permian - all extinct.
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FIGURE 12-52 Some common Paleozoic
blastoids
.
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Class Asteroidea - StarfishStarfish are star-shaped echinoderms with five arms.
About 430 genera are known. Geologic range: Ordovician to Holocene.
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FIGURE 12-48
Partially
dissected starfish
showing elements
of the water vascular system and other organs.
108Slide109
Class Ophiuroidea – Brittle StarsBrittle stars have 5 arms, like starfish, but the arms are thin and serpent-like.
About 325 genera are known. Geologic range: Ordovician to Holocene.
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FIGURE 12-47 Representative living echinoderms.
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Class EchninodeaSand dollars and sea urchins
Echinoids are disk-shaped, biscuit-shaped, or globular. Viewed from above, they may be circular or somewhat irregular in shape, but with a five-part symmetry. About 765 genera are known. Geologic range: Ordovician to Holocene.
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110Slide111
Class HolothuroideaSea cucumbers
Soft-bodied echinoderms resembling cucumbers. They have microscopic hard parts called sclerites in various shapes resembling hooks, wheels and anchors. About 200 genera are known.
Geologic range
: Middle Cambrian?; Middle Ordovician to Holocene
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111Slide112
Class EdrioasteroideaEdrioasteroids
A group that was probably ancestral to starfish and sea urchins.Globular, discoidal, or cylindrical tests (shells), many of which had concave surfaces. Geologic range: Early Cambrian to Middle Pennsylvanian.
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112Slide113
Class CystoideaCystoids
This primitive group had a calyx attached to the seafloor by a stem (like crinoids and blastoids). Distinctive patterns of pores on the plates of the calyx. Geologic range: Cambrian to Late Devonian. Most common during Ordovician and Silurian.
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113Slide114
The Echinoderm-Backbone Connection Echinoderms are
closely related to chordates (the group that includes the vertebrates). The early cell division, embryonic development, and larvae
of echinoderms resemble those of chordates, and are different from those of other invertebrates.
Biochemistry
of echinoderms is also similar to that of chordates (chemical similarities associated with muscle activity and chemistry of oxygen-carrying pigments in the blood).
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114Slide115
GraptolitesChief characteristics:
Organic (chitinous) skeletons consisting of rows or lines of small tubes or cups, called thecae. Tubes or cups branch off a main cord or tube called a stem or stipe.
Stipes may consist of one, two, or many branches.
Entire colony called a
rhabdosome
.
A filament at the lower end of the rhabdosome is called a
nema
.
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115Slide116
GraptolitesMost graptolites are found flattened and carbonized in black shales and mudstones.
Geologic range: Cambrian to Mississippian. (Most abundant during Ordovician and Silurian.)Some living organisms which may be surviving descendants (living fossils) have been recovered in 1989 in the South Pacific and later in Bermuda. Mode of Life
: Planktonic (colonies attached to floats).
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116Slide117
FIGURE 12-94 Five major mass extinction episodes. Source: After J. J. Sepkowski Jr., 1994, Geotimes,39(3): 15–17. FIGURE 12-2
Geologic time scale across the Proterozoic–Cambrian boundary. Source: Harold Levin. FIGURE 12-1 Tiny shell-bearing fossils from the LatePrecambrian and Early Cambrian in Siberia
. Source: Matthews, C., and
Missarzhevsky
, V., 1975, Small
shelly
fossils of late Precambrian and Early Cambrian age. Journal of the Geol. Society of London, 131:289-304.
FIGURE 12-5 The Burgess Shale fauna
.
Source: Harold Levin.
FIGURE 12-13 Reconstruction of
Pikaia, the earliest known
member of our own phylum,
Chordata
.
Source: Harold Levin.
FIGURE 12-8
Anomalocaris
, “invertebrate equivalent
of the
dinosaurs
.”
Source: Harold Levin.
FIGURE
12-10
Marrella
, the most elegant and
common arthropod
in the Burgess Shale fauna
.
Source: Harold Levin.
FIGURE
12-11 The early Cambrian Burgess Shale
fossil
Hallucigenia
.
Source: Harold Levin.
FIGURE 12-17
Archaeocyathid
skeleton. Source: Harold Levin.
FIGURE 12-18 Early Paleozoic sponges. Source: Harold Levin.
FIGURE 12-23 Common cnidarians
.
Source: Harold Levin.
FIGURE
12-24
Medusaandpolypformsin
cnidarians
.
Source: Harold Levin.
FIGURE
12-28
Dwelling positions of articulate
and inarticulate
brachiopods
.
Source: Harold Levin.
FIGURE 12-35 The
monoplacophoran
Pilina
.
Source: Harold Levin.
FIGURE 12-34 A common
placophoran
, the
Atlantic Coast
chiton
.
Source: Harold Levin.
FIGURE 12-36 Paleozoic bivalves. Source: Harold Levin.
FIGURE 12-39 Cephalopod suture patterns
.
Source: Harold Levin.
FIGURE
12-42
Trilobites
. Source: Harold Levin.
FIGURE
12-46
Mischoptera
, a
Pennsylvanian-age dragonfly.
Source: Harold Levin.
FIGURE 12-47 Representative living echinoderms
.
Source: Harold Levin.
FIGURE 12-55
Crinoid
in living position on
the seafloor.
Source: Harold Levin.
FIGURE 12-52 Some common Paleozoic
blastoids
.
Source: Harold Levin.
FIGURE 12-48 (A) Partially dissected starfish
showing elements
of the water vascular system and other organs
.
Source: Harold Levin.
IMAGE CREDITS
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117